Reptiles of Distrito Federal [Brazil] Related checklists:  Brazil Bibliography of the reptiles of Distrito Federal Taxonomic history of the reptiles of Distrito Federal Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Amphisbaena (Neotropical Worm Lizards)   Amphisbaena alba Common Red Worm Lizard Colli & Zamboni 1999 Costa, Guedes & Bernils 2021 Mott 2006 Pena Correa & Meneses 2020   Amphisbaena anaemariae Serra da Mesa Worm Lizard Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Gomes & Maciel 2012 Mott & Vieites 2009 Pena Correa & Meneses 2020 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019   Amphisbaena mertensii Sao Paulo Worm Lizard Costa, Guedes & Bernils 2021 Pena Correa & Meneses 2020   Amphisbaena talisiae Serra da Pitomba Worm Lizard Campos Nogueira 2001 [as A. mensae] Costa, Grabosky & Zaher 2019 Costa, Guedes & Bernils 2021 Pena Correa & Meneses 2020   Amphisbaena vermicularis Vermiculate Worm Lizard Costa, Guedes & Bernils 2021 Pena Correa & Meneses 2020 Genus Leposternon (South American Worm Lizards)   Leposternon infraorbitale Central Brazilian Worm Lizard Costa, Guedes & Bernils 2021 Ribeiro, Vaz-Silva & Santos 2008   Leposternon microcephalum Small-headed Worm Lizard Campos Nogueira 2001 Costa, Guedes & Bernils 2021

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Alligatoridae (Alligators and Caimans) Genus Caiman (Typical Caimans)   Caiman crocodilus Common Caiman Costa, Guedes & Bernils 2021 Genus Paleosuchus (Dwarf Caimans)   Paleosuchus palpebrosus Cayenne Dwarf Caiman Costa, Guedes & Bernils 2021

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Ophiodes (South American Glass Lizards)   Ophiodes striatus Dark-striped Glass Lizard Costa, Guedes & Bernils 2021  Family Anolidae (Anoles) Genus Norops (Brilliant Anoles)   Norops brasiliensis Brazilian Golden-scaled Anole Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Ribeiro-Junior 2015   Norops meridionalis Paraguayan Anole Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Ledo & Colli 2016 Ribeiro-Junior 2015  Family Gymnophthalmidae (Scaly-tongued Spectacled Tejus) Genus Bachia (Earless Tejus)   Bachia bresslaui Sao Paulo Earless Teju Colli, Zatz & Cunha 1998 Costa, Guedes & Bernils 2021 Rodrigues, Camacho, Nunes, Recoder, Teixeira, Valdujo, Ghellere, Mott & Nogueira 2008 Rodrigues, Pellegrino, Dixo, Verdade, Pavan, Argolo & Sites 2007 Teixeira, Recoder, Camacho, Sena, Navas & Rodrigues 2013 Genus Cercosaura (Eastern Shade Tejus)   Cercosaura olivacea East Brazilian Shade Teju Costa, Guedes & Bernils 2021 Ledo & Colli 2016 [as ocellata] Noronha-Oliveira, Carvalho, Freitas & Faria 2012 [as ocellata]   Cercosaura schreibersii South Brazilian Shade Teju Costa, Guedes & Bernils 2021 Ledo & Colli 2016 Ribeiro-Junior & Amaral 2017 Genus Colobosaura (Short-thumbed Spectacled Tejus)   Colobosaura modesta Morro da Garcza Spectacled Teju Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Freire, Jorge & Ribeiro 2012 Ribeiro-Junior & Amaral 2017 Genus Micrablepharus (Snake-eyed Glossy Tejus)   Micrablepharus atticolus Cerrado Snake-eyed Glossy Teju Costa, Guedes & Bernils 2021 Ledo & Colli 2016 Ribeiro-Junior & Amaral 2017 Rodrigues 1996   Micrablepharus maximiliani Blue-tailed Snake-eyed Glossy Teju Costa, Guedes & Bernils 2021 Moura, Dayrell & Avelar Sao Pedro 2010 Ribeiro-Junior & Amaral 2017  Family Hoplocercidae (Club-tailed Iguanas) Genus Hoplocercus (Eastern Club-tailed Iguana)   Hoplocercus spinosus Eastern Club-tailed Iguana Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015  Family Leiosauridae (Smooth Iguanas) Genus Enyalius (Fat-headed Anoles)   Enyalius bilineatus Two-lined Fat-headed Anole Costa, Guedes & Bernils 2021   Enyalius capetinga Cerrado Fat-headed Anole Breitman, Domingos, Bagley, Wiederhecker, Ferrari, Cavalcante, Pereira, Abreu, De-Lima, Morais, Prette, Silva, Mello, Carvalho, Lima, Silva, Matias, Carvalho, Pantoja, Gomes, Paschoaletto, Rodrigues, Talarico, Barreto-Lima & Colli 2018 Campos Nogueira 2001 [as bilineatus] Costa, Guedes & Bernils 2021 Ledo & Colli 2016 [as bilineatus] Rodrigues, Bertolotto, Amaro, Yonenaga-Yassuda, Freire & Pellegrino 2014 [as bilineatus] Rodrigues, Freitas, Silva & Bertolotto 2006 [as bilineatus] Sales, Ribeiro & Freire 2015 [as bilineatus]  Family Polychrotidae (Monkey Lizards) Genus Polychrus (Monkey Lizards)   Polychrus acutirostris Sharp-nosed Monkey Lizard Avila-Pires 1995 Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015  Family Scincidae (Skinks) Genus Aspronema (South American White-striped Skinks)   Aspronema dorsivittatum South American White-striped Skink Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Hedges & Conn 2012 Ledo & Colli 2016 Miralles & Carranza 2010 Genus Copeoglossum (Neotropical Spotted Skinks)   Copeoglossum nigropunctatum Black-spotted Skink Costa, Guedes & Bernils 2021 Hedges & Conn 2012 Ledo & Colli 2016 Miralles & Carranza 2010 Miralles, Barrio-Amoros, Rivas & Chaparro-Auza 2006 Ribeiro-Junior & Amaral 2016 Genus Manciola (South American Small-handed Skink)   Manciola guaporicola South American Small-handed Skink Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Ribeiro-Junior & Amaral 2016 Genus Notomabuya (Southern Neotropical Skink)   Notomabuya frenata Southern Neotropical Skink Caldeira Costa, Dias Fernandes, Vrcibradic & Neves Feio 2008 Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Ledo & Colli 2016 Ribeiro-Junior & Amaral 2016  Family Teiidae (Greater Tejus) Genus Ameiva (South American Ameivas)   Ameiva ameiva South American Giant Ameiva Colli 1991 Costa, Guedes & Bernils 2021 Ledo & Colli 2016 Ribeiro-Junior & Amaral 2016 Genus Ameivula (Eastern South American Whiptails)   Ameivula ocellifera Brazilian Ocellated Whiptail Aparicio, Avila-Pires, Moravec & Nogueira 2019 Costa, Guedes & Bernils 2021 Franca, Mesquita, Noguiera & Araujo 2008 Mesquita & Colli 2003 Genus Kentropyx (Spurred Whiptails)   Kentropyx calcarata Striped Forest Whiptail Costa, Guedes & Bernils 2021   Kentropyx paulensis Sao Paulo Whiptail Costa, Guedes & Bernils 2021 Gallagher & Dixon 1992 Genus Salvator (Southern Tejus)   Salvator duseni Parana Teju Campos, Oda, Custodio, Koppe & Dartora 2011 Colli, Peres & Cunha 1998 [as rufescens] Costa, Guedes & Bernils 2021   Salvator merianae Southern Black-and-White Teju Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Genus Tupinambis (Northern Tejus)   Tupinambis quadrilineatus Four-striped Teju Costa, Guedes & Bernils 2021 Guimaraes, Figueiredo & Salmito 2007 Ribeiro-Junior & Amaral 2016 Silva, Ribeiro-Junior & Avila-Pires 2018 Silveira 2009  Family Tropiduridae (Rough-tailed Iguanas) Genus Tropidurus (Eastern Keel-tailed Iguanas)   Tropidurus itambere Sorocaba Keel-tailed Iguana Costa, Guedes & Bernils 2021 Ledo & Colli 2016   Tropidurus oreadicus Cerrado Keel-tailed Iguana Carvalho 2013 Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015   Tropidurus torquatus Collared Keel-tailed Iguana Costa, Guedes & Bernils 2021 Ledo & Colli 2016 Vieira, Wiederhecker, Colli & Bao 2001

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Anomalepididae (Neotropical Blind Snakes) Genus Liotyphlops (Smooth Blind Snakes)   Liotyphlops ternetzii Paraguayan Blind Snake Costa, Guedes & Bernils 2021 Cruz-Centeno, Swaya & Germano 2010 Dixon & Kofron 1984 Franca & Araujo 2006 McDiarmid, Campbell & Toure 1999 Genus Boa (Red-tailed Boas)   Boa constrictor South American Red-tailed Boa Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Epicrates (Rainbow Boas)   Epicrates crassus Paraguayan Rainbow Boa Costa, Guedes & Bernils 2021 Franca & Araujo 2006 [as cenchria] Peters & Orejas-Miranda 1970 [as cenchria polylepis] Genus Eunectes (Anacondas)   Eunectes murinus Green Anaconda Costa, Guedes & Bernils 2021 Franca & Araujo 2006  Family Colubridae (Typical Snakes or Colubrids) Genus Adelphostigma (Twin-spotted Leaf-litter Snakes)   Adelphostigma occipitalis Bahia Leaf-litter Snake Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Apostolepis (Black-tailed Blackheads)   Apostolepis albicollaris White-collared Blackhead Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes, Paredero, Tiutenko & Loebmann 2022 Franca & Araujo 2006 Lema & Renner 2012 Lema 2002 Nogueira, Barbo & Ferrarezzi 2012 Wallach, Williams & Boundy 2014   Apostolepis assimilis Common Red Blackhead Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Ferrarezzi, Barbo & Albuquerque 2005 Franca & Araujo 2006 Harvey 1999 Lema & Renner 2011 Lema & Renner 2012 Lema 2002 Nogueira, Barbo & Ferrarezzi 2012   Apostolepis flavotorquata Yellow-collared Blackhead Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Lema & Renner 2005 Lima Silveira 2014 Wallach, Williams & Boundy 2014   Apostolepis goiasensis Goias Blackhead Costa, Guedes & Bernils 2021 Maffei & Campos Nogueira 2021   Apostolepis sanctaeritae Tocantins Blackhead Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Ferrarezzi, Barbo & Albuquerque 2005 [as ammodites] Lema & Renner 2007 [as ammodites] Nogueira, Barbo & Ferrarezzi 2012 [as ammodites] Wallach, Williams & Boundy 2014 [as ammodites] Genus Atractus (Spindle Snakes)   Atractus pantostictus Sao Paulo Spindle Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Passos, Fernandes, Bernils & Moura-Leite 2010 Genus Boiruna (Southern Snake Eaters)   Boiruna maculata South American Spotted Snake Eater Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Chironius (Sipos)   Chironius brazili Southeast Brazilian Sipo Costa, Guedes & Bernils 2021 Hamdan & Fernandes 2015   Chironius exoletus Green Sipo Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993 Franca & Araujo 2006   Chironius flavolineatus Yellow-striped Sipo Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Hamdan & Fernandes 2015 Hamdan, Scali & Fernandes 2014 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010   Chironius quadricarinatus Paraguayan Sipo Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010 Genus Clelia (Northern Snake Eaters)   Clelia plumbea Plumbeous Snake Eater Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Zaher 1996 Genus Dipsas (Neotropical Snail Eaters)   Dipsas mikanii Brazilian White-collared Snail Eater Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Drymarchon (Indigo Snakes and Cribos)   Drymarchon corais Yellow-tailed Cribo Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Drymoluber (Glossy Racers)   Drymoluber brazili Southern Glossy Racer Caldeira-Costa, Ribeiro-Moura & Neves-Feio 2013 Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Freitas, Franca & Verissimo 2012 Genus Dryophylax (Northern Mock Vipers)   Dryophylax hypoconia Southern Mock Viper Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Erythrolamprus (Neotropical Smooth Snakes)   Erythrolamprus aesculapii Southern Mock Coral Snake Costa, Guedes & Bernils 2021 Curcio, Scali & Rodrigues 2015 Franca & Araujo 2006   Erythrolamprus almadensis Almada Smooth Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Erythrolamprus macrosomus Southern Royal Smooth Snake Costa, Guedes & Bernils 2021   Erythrolamprus maryellenae Goias Smooth Snake Costa, Guedes & Bernils 2021 Dixon 1985 Franca & Araujo 2006   Erythrolamprus poecilogyrus Yellow-bellied Smooth Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Erythrolamprus reginae Northern Royal Smooth Snake Franca & Araujo 2006 Genus Gomesophis (Brazilian Burrowing Snake)   Gomesophis brasiliensis Brazilian Burrowing Snake Castellari-Gonzalez, Prudente & Franco 2014 Costa, Guedes & Bernils 2021 Fortes, Lucas & Caldart 2010 Franca & Araujo 2006 Menezes, Fiorillo, Franco & Feio 2017 Prudente & Brandao 1998 Genus Helicops (South American Keelbacks)   Helicops angulatus Brown-banded Keelback Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Roberto, Ribeiro, Sousa-Delfino & Oliveira-Almeida 2009   Helicops infrataeniatus Surinam Keelback Costa, Guedes & Bernils 2021   Helicops leopardinus Leopard Keelback Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021   Helicops modestus Brazilian Olive Keelback Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Freitas & Silva 2011 Genus Leptodeira (Typical Cat-eyed Snakes)   Leptodeira annulata South American Banded Cat-eyed Snake Barrio-Amoros 2019 Costa, Guedes & Bernils 2021 Duellman 1958 Franca & Araujo 2006 Genus Lygophis (South American Striped Ground Snakes)   Lygophis meridionalis Paraguayan Striped Ground Snake Castro & Oliveira 2017 Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Lygophis paucidens Cerrado Striped Ground Snake Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Serrano, Vieira-Alencar, Diaz-Ricaurte & Nogueira 2020 Genus Mesotes (Southern Mock Vipers)   Mesotes rutilus Sao Paulo Mock Viper Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Franco & Ferreira 2002 Genus Mussurana (Brazilian Snake Eaters)   Mussurana quimi South Brazilian Snake Eater Costa, Guedes & Bernils 2021 Entiauspe-Neto, Abegg, Quintela & Loebmann 2017 Franca & Araujo 2006 Franco, Marques & Puorto 1997 Giraudo & Scrocchi 2002 Silveira & Cotta 2006 Genus Oxybelis (Neotropical Vine Snakes)   Oxybelis aeneus Brown Vine Snake Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Oxyrhopus (Calico Snakes)   Oxyrhopus guibei Parana Calico Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Silveira 2008   Oxyrhopus petolarius Banded Calico Snake Costa, Guedes & Bernils 2021 Franca, Barros & Horta 2010   Oxyrhopus rhombifer Rhombic Calico Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Oxyrhopus trigeminus Northern Brazilian Calico Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Peters & Orejas-Miranda 1970 Genus Palusophis (Eastern Lizard Eater)   Palusophis bifossatus Eastern Lizard Eater Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Montingelli 2009 Montingelli, Grazziotin, Battilana, Murphy, Zhang & Zaher 2019 Genus Phalotris (South American Collared Snakes)   Phalotris nasutus Snouted Collared Snake Costa, Guedes & Bernils 2021 Ferrarezzi 1993 Ferreira, Wang & Himmelstein 2005 Franca & Araujo 2006 Lema & Renner 2012 Lema 2002 Genus Philodryas (South American Bush Racers)   Philodryas aestiva Eastern Brazilian Green Bush Racer Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Philodryas nattereri Paraguay Bush Racer Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Smith, Scott, Cacciali, Atkinson & Pheasey 2013   Philodryas olfersii Common Green Bush Racer Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Philodryas psammophidea South American Striped Racer Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Franca 2003 Genus Phimophis (South American Shovel-nosed Snakes)   Phimophis guerini Argentine Shovel-nosed Snake Costa, Guedes & Bernils 2021 Filho, Santana, Vieira, Alves, Montenegro & Freitas 2012 Franca & Araujo 2006 Genus Pseudablabes (Reticulated Bush Racers)   Pseudablabes agassizii Burrowing Night Snake Carvalho-Junior, Schettino Canelas & Cordeiro-Malta 2006 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Souza Filho & Plombon 2014   Pseudablabes patagoniensis Patagonian Bush Racer Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Gouveia, Novelli, Vieira & Sousa 2017 Genus Pseudoboa (Neotropical Scarlet Snakes)   Pseudoboa nigra Blackish Scarlet Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Rhachidelus (Brazilian Bird Snake)   Rhachidelus brazili Brazilian Bird Snake Arruda, Avila, Morais & Kawashito-Ribeiro 2011 Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Lima Silveira 2004 Genus Simophis (South American False Coral Snake)   Simophis rhinostomus South American False Coral Snake Cacciali, Scott, Gunther, Sawaya, Brusquetti & Bauer 2009 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Spilotes (Chicken Snakes)   Spilotes pullatus Chicken Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Tantilla (American Centipede Snakes)   Tantilla melanocephala South American Black-headed Centipede Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Genus Xenodon (Northern Mock Lanceheads and South American Hognose Snakes)   Xenodon merremii Toad-eating Mock Lancehead Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Xenodon nattereri Banded Hognose Snake Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Lima Silveira, Ribeiro Costa & Salles 2004 Genus Xenopholis (South American Flat-headed Snakes)   Xenopholis undulatus Southern Flat-headed Snake Borges-Nojosa & Cassiano Lima 2009 Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Araujo 2006  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Micrurus (American Coral Snakes)   Micrurus carvalhoi Brazilian Ribbon Coral Snake Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Araujo 2006 [as M. lemniscatus] Pires, Silva, Feitosa, Prudente, Filho & Zaher 2014   Micrurus frontalis Cerrado Coral Snake Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Araujo 2006 Silva & Sites 1999  Family Viperidae (Vipers) Genus Bothrops (American Lanceheads)   Bothrops alternatus Urutu Carrasco, Grazziotin, Farfan, Koch, Ochoa, Scrocchi, Leynaud & Chaparro 2019   Bothrops itapetiningae Itapetininga Lancehead Carrasco, Grazziotin, Farfan, Koch, Ochoa, Scrocchi, Leynaud & Chaparro 2019 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Araujo 2006   Bothrops lutzi Cerrado Lancehead Costa, Guedes & Bernils 2021   Bothrops marmoratus Marbled Lancehead Carrasco, Grazziotin, Farfan, Koch, Ochoa, Scrocchi, Leynaud & Chaparro 2019 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018   Bothrops moojeni Brazilian Lancehead Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 Franca & Araujo 2006 Nogueira, Sawaya & Martins 2003   Bothrops neuwiedi Eastern Brazilian Lancehead Costa, Guedes & Bernils 2021 Franca & Araujo 2006   Bothrops pauloensis Black-faced Lancehead Costa, Guedes & Bernils 2021 Genus Crotalus (Typical Rattlesnakes)   Crotalus durissus South American Rattlesnake Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Araujo 2006 Nascimento, Avila Pires & Cunha 1988

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Chelidae (Austro-American Side-necked Turtles) Genus Acanthochelys (South American Side-necked Swamp Turtles)   Acanthochelys spixii Black Spiny-necked Swamp Turtle Costa, Guedes & Bernils 2021 Genus Mesoclemmys (Hump-backed Toad-headed Turtles)   Mesoclemmys vanderhaegei Tobati Toad-headed Turtle Costa, Guedes & Bernils 2021 Lima-Silveira 2009 Maffei, Nascimento, Moya & Donatelli 2016 Marques, Boehm, Brito, Cabrera & Verdade 2014 Genus Phrynops (Lesser Toad-headed Turtles)   Phrynops geoffroanus Brazilian Toad-headed Turtle Costa, Guedes & Bernils 2021  Family Testudinidae (Tortoises) Genus Chelonoidis (South American Tortoises)   Chelonoidis carbonarius South American Red-footed Tortoise Costa, Guedes & Bernils 2021