Reptiles of Botswana    Bibliography of the reptiles of Botswana Taxonomic history of the reptiles of Botswana Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Dalophia (Blunt-tailed Worm Lizards)   Dalophia longicauda Long-tailed Worm Lizard Auerbach 1987 Barts & Haacke 1997 Broadley, Gans & Visser 1976 Gans 2005   Dalophia pistillum Zambezi Blunt-tailed Worm Lizard Auerbach 1987 Barts & Haacke 1997 Broadley, Gans & Visser 1976 Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis anchietae Mossamedes Spade-snouted Worm Lizard Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley, Gans & Visser 1976 Kennedy, Marais, Bauer, Lewis & Thies 2012   Monopeltis capensis Cape Spade-snouted Worm Lizard Auerbach 1987 Branch 1998 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group A"] Gans 2005   Monopeltis infuscata Dusky Spade-snouted Worm Lizard Branch 1998 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group B"] Gans 2005   Monopeltis leonhardi Kalahari Spade-snouted Worm Lizard Auerbach 1987 Branch 1998 Broadley, Gans & Visser 1976 Gans 2005   Monopeltis mauricei Botswana Spade-snouted Worm Lizard Auerbach 1987 Branch 1998 Broadley 1992 Gans 2005   Monopeltis sphenorhynchus Slender Spade-snouted Worm Lizard Broadley, Gans & Visser 1976 Gans 2005 Genus Zygaspis (Purple Round-headed Worm Lizards)   Zygaspis nigra Black Round-headed Worm Lizard Auerbach 1987 Barts & Haacke 1997   Zygaspis quadrifrons Kalahari Round-headed Worm Lizard Barts & Haacke 1997 Broadley & Broadley 1997 Gans 2005 Kennedy, Marais, Bauer, Lewis & Thies 2012

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Auerbach 1987 King & Burke 1989 Wermuth & Mertens 1977

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus atricollis Black-necked Ridgeback Agama Auerbach 1987 Barts & Haacke 1997 Branch 1998 Wagner, Butler, Ceriaco & Bauer 2021 Wagner, Greenbaum & Bauer 2012 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018 Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Auerbach 1987 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012 Wermuth 1967   Agama armata Zambezi Spiny Agama Auerbach 1987 Barts & Haacke 1997 Branch 1998   Agama atra Southern Rock Agama Auerbach 1987 Wermuth 1967   Agama kirkii Zambesi Rock Agama Auerbach 1987 Branch 1998 Broadley 2002   * Agama makarikarica Makgadikgadi Spiny Agama Auerbach 1987 Branch 1998 Wermuth 1967  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Auerbach 1987 Barts & Haacke 1997 Branch 1998 Glaw 2015 Kennedy, Marais, Bauer, Lewis & Thies 2012 Klaver & Boehme 1997 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Cordylus (Girdled Lizards)   Cordylus jonesii Limpopo Girdled Lizard Auerbach 1987 Branch 1998 Broadley & Branch 2002 Wermuth 1968   Cordylus vittifer Transvaal Girdled Lizard Auerbach 1987 Bates & Broadley 2012 Wermuth 1968 Genus Platysaurus (Flat Lizards)   Platysaurus intermedius Common Flat Lizard Auerbach 1987 Branch 1998 Broadley 1978  Family Gekkonidae (Typical Geckos) Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus angulifer Giant Ground Gecko Auerbach 1987 Heinz, Brennan, Jackman & Bauer 2021   Chondrodactylus bibronii Tubercled Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Heinz, Brennan, Jackman & Bauer 2021   Chondrodactylus laevigatus Karas Thick-toed Gecko Heinz, Brennan, Jackman & Bauer 2021   Chondrodactylus turneri Zambezi Thick-toed Gecko Auerbach 1987 [as Pachydactylus bibronii] Heinz, Brennan, Jackman & Bauer 2021 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus mabouia Cosmopolitan House Gecko Auerbach 1987 Branch 1998 Broadley 1977 Loveridge 1947 Loveridge 1957 Genus Homopholis (African Velvet Geckos)   Homopholis arnoldi Mahalapye Velvet Gecko Auerbach 1987 [as wahlbergii] Branch 1998 [as wahlbergii] Broadley, Jackman & Bauer 2014 Loveridge 1944 Loveridge 1947 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus angolensis Angolan Dwarf Gecko Branch 1998 Loveridge 1947 Loveridge 1957 Wermuth 1965   Lygodactylus bradfieldi Namibian Dwarf Gecko Auerbach 1987 Branch 1998   Lygodactylus capensis Cape Dwarf Gecko Auerbach 1987 Barts & Haacke 1997 Kennedy, Marais, Bauer, Lewis & Thies 2012 Loveridge 1947   Lygodactylus chobiensis Chobe Dwarf Gecko Auerbach 1987 Loveridge 1947 Wermuth 1965 Genus Pachydactylus (Typical Thick-toed Geckos)   Pachydactylus capensis Common Cape Thick-toed Gecko Auerbach 1987 Broadley 1977   Pachydactylus punctatus Speckled Thick-toed Gecko Auerbach 1987 Branch 1998   Pachydactylus tigrinus Tiger Thick-toed Gecko Auerbach 1987 Branch 1998 Broadley 1977   * Pachydactylus tsodiloensis Tsodilo Thick-toed Gecko Barts & Haacke 1997 Bauer, Lamb & Branch 2006   Pachydactylus wahlbergii Kalahari Ground Gecko Auerbach 1987 Barts & Haacke 1997 Branch 1998 Genus Ptenopus (African Barking Geckos)   Ptenopus garrulus Common Barking Gecko Auerbach 1987 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012  Family Gerrhosauridae (Plated Lizards) Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus auritus Kalahari Plated Lizard Auerbach 1987 Barts & Haacke 1997 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 Wermuth 1968   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Auerbach 1987   Gerrhosaurus intermedius Eastern Black-lined Plated Lizard Auerbach 1987 [as n. nigrolineatus] Bates, Tolley, Edwards, Davids, Silva & Branch 2013 [as nigrolineatus] Genus Matobosaurus (Giant Plated Lizards)   Matobosaurus validus Eastern Giant Plated Lizard Auerbach 1987 Bates, Tolley, Edwards, Davids, Silva & Branch 2013  Family Lacertidae (Typical Lizards) Genus Heliobolus (Bushveld Lizards)   Heliobolus lugubris Southern African Bushveld Lizard Auerbach 1987 Barts & Haacke 1997 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012 Szczerbak 1975 Genus Ichnotropis (Rough-scaled Lizards)   Ichnotropis capensis Southern Rough-scaled Lizard Auerbach 1987 Barts & Haacke 1997 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012   Ichnotropis grandiceps Caprivi Rough-scaled Lizard Auerbach 1987 Barts & Haacke 1997 Branch 1998 Genus Meroles (Southwest African Desert Lizards)   Meroles squamulosus Rough-scaled Desert Lizard Auerbach 1987 Barts & Haacke 1997 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012 Loveridge 1957   Meroles suborbitalis Spotted Desert Lizard Auerbach 1987 Genus Nucras (Sandveld Lizards)   Nucras holubi Southern Sandveld Lizard Branch 1998   Nucras intertexta Spotted Sandveld Lizard Auerbach 1987 Broadley 1972   Nucras ornata Ornate Sandveld Lizard Auerbach 1987 Broadley 1972   Nucras tessellata Western Sandveld Lizard Auerbach 1987 Branch 1998 Broadley 1972 Genus Pedioplanis (African Sand Lizards)   Pedioplanis lineoocellata Spotted Sand Lizard Auerbach 1987 Branch 1998 Szczerbak 1975   Pedioplanis namaquensis Namaqua Sand Lizard Auerbach 1987 Branch 1998 Kennedy, Marais, Bauer, Lewis & Thies 2012 Szczerbak 1975  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias gariepensis Gariep Blind Legless Skink Auerbach 1987 Branch 1998   Acontias kgalagadi Kgalagadi Legless Skink Auerbach 1987 [as Typhlosaurus lineatus] Barts & Haacke 1997 [as Typhlosaurus l. lineatus] Branch 1998 [as Typhlosaurus l. lineatus] Broadley 1968 [as Typhlosaurus lineatus] Conradie & Bourquin 2013 Kennedy, Marais, Bauer, Lewis & Thies 2012   Acontias occidentalis Western Black Legless Skink Auerbach 1987 Broadley & Greer 1969 Griffin 2003 Genus Mochlus (African Supple Skinks)   Mochlus sundevallii Common Savanna Supple Skink Auerbach 1987 Barts & Haacke 1997 Broadley 1966 Kennedy, Marais, Bauer, Lewis & Thies 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis maculicollus Spotted-neck Snake-eyed Skink Jacobsen & Broadley 2000   Panaspis wahlbergii Savanna Snake-eyed Skink Auerbach 1987 Branch 1998 Genus Scelotes (Dwarf Burrowing Skinks)   Scelotes limpopoensis Limpopo Dwarf Burrowing Skink Branch 1998 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis capensis Cape Skink Auerbach 1987 Broadley 2000   Trachylepis damarana Damara Variable Skink Auerbach 1987 [as varia] Barts & Haacke 1997 [as varia] Kennedy, Marais, Bauer, Lewis & Thies 2012 [as varia] Portik, Bauer & Jackman 2010 [as varia] Weinell & Bauer 2018   Trachylepis margaritifer African Rainbow Skink Auerbach 1987 Broadley & Bauer 1998 Broadley 2000   Trachylepis occidentalis Western Three-striped Skink Auerbach 1987 Branch 1998 Broadley 2000   Trachylepis punctatissima Montane Speckled Skink Auerbach 1987 Branch 1998 Broadley 1977 Broadley 2000   Trachylepis punctulata Eastern Variegated Skink Auerbach 1987 Branch 1998 Broadley 1975 Broadley 2000   Trachylepis sparsa Karasburg Tree Skink Auerbach 1987 Branch 1998 Broadley 1977 Broadley 2000   Trachylepis spilogaster Namibian Tree Skink Auerbach 1987 Branch 1998 Broadley 1977 Broadley 2000   Trachylepis striata Eastern Striped Skink Auerbach 1987   Trachylepis wahlbergii Mopane Skink Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley 1977 Broadley 2000 Kennedy, Marais, Bauer, Lewis & Thies 2012 Genus Typhlacontias (Western Burrowing Skinks)   Typhlacontias rohani Kalahari Burrowing Skink Auerbach 1987 Branch 1998 Haacke 1997  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Barts & Haacke 1997 Bayless 2002   Varanus niloticus Nile Monitor Auerbach 1987 Bayless 2002 Lenz in Pianka, King & King (eds.) 2004

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis barotseensis Zambian Herald Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1983   Crotaphopeltis hotamboeia Common Herald Snake Auerbach 1987 Broadley & Blaylock 2013 Spawls & Graff 1989 Genus Dasypeltis (African Egg Eaters)   Dasypeltis scabra Common African Egg Eater Auerbach 1987 Bates & Broadley 2018 Broadley & Blaylock 2013 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Broadley & Blaylock 2013 Pietersen, Verburght & Davies 2021 Genus Limnophis (Striped Swamp Snakes)   Limnophis bangweolicus Eastern Striped Swamp Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1983 Chippaux & Jackson 2019 Conradie, Deepak, Keates & Gower 2020 Genus Meizodon (African Smooth Snakes)   Meizodon semiornatus Semi-ornate Smooth Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1983 Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1966 Broadley 1983 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013   Philothamnus ornatus Ornate Green Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1983   Philothamnus semivariegatus Variegated Bush Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1983 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus semiannulatus Southern Tiger Snake Auerbach 1987 Broadley & Blaylock 2013 Loveridge 1957 Genus Thelotornis (African Vine Snakes)   Thelotornis capensis Southern Vine Snake Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2002 Broadley 1983 Broadley 2001 Spawls & Branch 1995 Spawls & Graff 1989  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps scutatus Shield-nosed Coral Snake Auerbach 1987 Branch 1998 Broadley & Baldwin 2006 Broadley & Blaylock 2013 Broadley 1983 Wallach, Williams & Boundy 2014 Genus Dendroaspis (Mambas)   Dendroaspis polylepis Black Mamba Auerbach 1987 Barts & Haacke 1997 Broadley & Blaylock 2013 David & Ineich 1999 Haakansson & Madsen 1983 Kennedy, Marais, Bauer, Lewis & Thies 2012 Spawls & Branch 1995 Spawls & Graff 1989 Genus Elapsoidea (African Garter Snakes)   Elapsoidea boulengeri Zambezi Garter Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1971 Broadley 1983 Broadley 1998 David & Ineich 1999 [as E. semiannulata] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002   Elapsoidea sundevallii Southern African Garter Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1983 Broadley 1998 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja anchietae Southwest African Cobra Auerbach 1987 Bauer, Kennedy, Lewis, Thies & Gabadirwe 2009 Branch 1998 Broadley & Blaylock 2013 Broadley & Wuster 2004 Broadley 1968 Broadley 1983 Broadley 1995 Spawls & Branch 1995   Naja annulifera African Banded Cobra Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wuster 2004 Broadley 1968 Broadley 1983 Broadley 1995 Spawls & Branch 1995   Naja mossambica Mozambique Spitting Cobra Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley & Blaylock 2013 David & Ineich 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Naja nivea Yellow Cobra Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wuster 2004 Broadley 1968 Spawls & Branch 1995  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1971 Broadley 1983 Spawls & Graff 1989   Amblyodipsas mechowii Elongate Quill-snouted Snake Auerbach 1987 Barts & Haacke 1997 Broadley & Blaylock 2013 Broadley 1971 Broadley 1983 Spawls & Graff 1989   Amblyodipsas polylepis Common Purple-glossed Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1983 Spawls & Graff 1989 Wallach, Williams & Boundy 2014   Amblyodipsas ventrimaculata Kalahari Purple-glossed Snake Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1983 Spawls & Graff 1989 Genus Aparallactus (African Centipede Eaters)   Aparallactus capensis Cape Centipede Eater Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Loveridge 1957 Spawls & Graff 1989 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1991 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Spawls & Graff 1989 Wallach, Williams & Boundy 2014   Atractaspis congica Congo Stiletto Snake Branch 1998   Atractaspis duerdeni Beaked Stiletto Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1991 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Laurent 1950 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Auerbach 1987 [as fuliginosus] Bauer, Kennedy, Lewis, Thies & Gabadirwe 2009 [as fuliginosus] Broadley & Blaylock 2013   Boaedon fradei Moxico House Snake Hallermann, Ceriaco, Schmitz, Ernst, Conradie, Verburgt, Marques & Bauer 2020 Genus Dipsina (Dwarf Beaked Snake)   Dipsina multimaculata Dwarf Beaked Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Chirio & Ineich 1991 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley & Howell 1991 Hughes 2004 Genus Gracililima (Black File Snake)   Gracililima nyassae Black File Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1983 Lanza & Broadley 2014 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley & Hughes 2000 Broadley 1983 Genus Lamprophis (SubSaharan House Snakes)   Lamprophis aurora Aurora House Snake Broadley & Blaylock 2013 Broadley 1990 Genus Limaformosa (Venus File Snakes)   Limaformosa capensis Cape File Snake Auerbach 1987 Broadley & Blaylock 2013 Loveridge 1957 Spawls & Graff 1989 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1996 Spawls & Graff 1989 Genus Prosymna (Shovel-snout Snakes)   Prosymna angolensis Angolan Shovel-snout Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1983 Conradie, Keates, Baptista & Lobon-Rovira 2022   Prosymna bivittata Two-striped Shovel-snout Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1983   Prosymna lineata Lineolate Shovel-snout Auerbach 1987 Broadley & Blaylock 2013 Broadley 1980 Broadley 1983 Spawls & Graff 1989   Prosymna sundevallii Southern African Shovel-snout Auerbach 1987 Broadley & Blaylock 2013 Broadley 1980 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis angolensis Dwarf Sand Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1983 Broadley 2002   Psammophis brevirostris Short-snouted Grass Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 1977 Broadley 1983 Broadley 2002   Psammophis jallae Zimbabwe Sand Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1977 Broadley 2002   Psammophis mossambicus East African Olive Grass Snake Auerbach 1987 [as P. phillipsi] Barts & Haacke 1997 [as P. phillipsi] Bauer, Kennedy, Lewis, Thies & Gabadirwe 2009 Broadley & Blaylock 2013 [as P. phillipsii] Broadley 1977 [as P. phillipsi] Broadley 1983 [as P. phillipsii] Broadley 2002 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis notostictus Karoo Sand Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley 2002   Psammophis subtaeniatus Western Stripe-bellied Sand Snake Auerbach 1987 Barts & Haacke 1997 Broadley & Blaylock 2013 Broadley 1977 Broadley 1983 Broadley 2002 Kennedy, Marais, Bauer, Lewis & Thies 2012   Psammophis trinasalis Kalahari Sand Snake Auerbach 1987 Bauer, Kennedy, Lewis, Thies & Gabadirwe 2009 Branch 1998 Broadley & Blaylock 2013 Broadley 1977 Broadley 1983 Broadley 2002 Kennedy, Marais, Bauer, Lewis & Thies 2012 Spawls & Graff 1989 Genus Psammophylax (Skaapstekers)   Psammophylax tritaeniatus Southern Striped Skaapsteker Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1976 Keates, Conradie, Greenbaum & Edwards 2019   Psammophylax variabilis Grey-bellied Skaapsteker Auerbach 1987 Broadley & Blaylock 2013 Broadley & Howell 1991 Broadley 1976 Broadley 1983 Largen & Rasmussen 1993 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Auerbach 1987 Barts & Haacke 1997 Broadley & Blaylock 2013 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis rostratus Rufous Beaked Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops scutifrons Zambezi Thread Snake Auerbach 1987 Barts & Haacke 1997 Broadley & Blaylock 2013 Broadley & Broadley 1999 Broadley & Watson 1976 Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis longicauda Southern African Long-tailed Thread Snake Auerbach 1987 Broadley & Broadley 1999 Broadley & Watson 1976 Hahn 1980 McDiarmid, Campbell & Toure 1999  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python natalensis Southern Rock Python Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley & Blaylock 2013 Broadley & Howell 1991 Broadley 1983 Broadley 1984 Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops bibronii Southern African Blind Snake Auerbach 1987 Broadley & Blaylock 2013 Broadley & Wallach 2009 Spawls & Graff 1989 Wallach, Williams & Boundy 2014   Afrotyphlops mucruso Zambezi Giant Blind Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1983 Spawls, Howell, Drewes & Ashe 2002   Afrotyphlops schlegelii Variable Giant Blind Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1983 Spawls & Graff 1989 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops boylei Kalahari Beaked Blind Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1983   Rhinotyphlops lalandei Cape Blind Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1983 McDiarmid, Campbell & Toure 1999 Spawls & Graff 1989   Rhinotyphlops schinzi Black-spotted Beaked Blind Snake Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1983  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Auerbach 1987 Barts & Haacke 1997 Branch 1998 Broadley & Blaylock 2013 Broadley 1983 David & Ineich 1999 Kennedy, Marais, Bauer, Lewis & Thies 2012 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995   Bitis caudalis Horned Adder Auerbach 1987 Broadley & Blaylock 2013 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus rhombeatus Rhombic Night Adder Auerbach 1987 Branch 1998 Broadley & Blaylock 2013 Broadley 1983 David & Ineich 1999 Spawls & Branch 1995

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa subrufa Common Helmeted Turtle Auerbach 1987 Boycott & Bourquin 2008 Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios bechuanicus Okavango Mud Turtle Auerbach 1987 Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios rhodesianus Zambian Mud Turtle Auerbach 1987 Broadley & Boycott 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios sinuatus East African Serrated Mud Turtle Auerbach 1987 Broadley & Boycott 2009 Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios subniger East African Black Mud Turtle Auerbach 1987 Branch 1998 Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys lobatsiana Lobatse Hinge-back Tortoise Branch 1998 Broadley 1993 Fritz & Havas 2006 Ihlow, Huyssteen, Vamberger, Cory-Toussaint, Hofmeyr & Fritz 2020 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Kinixys spekii Eastern Savanna Hinge-back Tortoise Auerbach 1987 [as belliana] Hailey, Broadley, Boycott & Spawls 2021 Ihlow, Farooq, Gvozdik, Hofmeyr, Conradie, Campbell, Harvey, Verburgt & Fritz 2019 Iverson 1992 King & Burke 1989 [as belliana] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wermuth & Mertens 1977 [as belliana] Genus Psammobates (South African Star Tortoises)   Psammobates oculiferus African Serrated Star Tortoise Auerbach 1987 Barts & Haacke 1997 Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wermuth & Mertens 1977 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Auerbach 1987 Barts & Haacke 1997 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021