Reptiles of Gambia    Bibliography of the reptiles of Gambia Taxonomic history of the reptiles of Gambia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca feae Guinea-Bissau Worm Lizard Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gans 1987 Gans 2005 Pauwels & Meirte 1996

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Barnett, Emms & Santoni 2001 [as niloticus] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as niloticus] Fuchs, Mertens & Wermuth 1974 Gruschwitz, Lenz & Boehme 1991 [as niloticus] Haakansson 1981 [as niloticus] Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   Mecistops cataphractus West African Slender-snouted Crocodile Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Shirley, Carr, Nestler, Vliet & Brochu 2018 Trape, Trape & Chirio 2012 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama bocourti Casamance Agama The Reptile Database (viewed 10th April, 2011)   Agama picticauda Ada Foah Agama Barnett, Emms & Santoni 2001 [as agama] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as agama] Gruschwitz, Lenz & Boehme 1991 [as agama] Haakansson 1981 [as agama] Pauwels & Meirte 1996 [as agama] Trape, Trape & Chirio 2012 [as agama]   Agama weidholzi Senegal Agama Boehme 2005 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Wagner & Bauer 2011 Wagner, Ineich, Leache, Wilms, Trape, Boehme & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo gracilis Graceful Chameleon Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Glaw 2015 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Klaver & Boehme 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Glaw 2015 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Tilbury 2010 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 [as brookii] Haakansson 1981 Loveridge 1947 Pauwels & Meirte 1996 [as brookii] Trape, Trape & Chirio 2012 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Trape, Trape & Chirio 2012  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola parvicarinata Small-keeled Gecko Trape, Trape & Chirio 2012   Tarentola senegambiae Senegambian Wall Gecko Barnett, Emms & Santoni 2001 [as ephippiata] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as ephippiata] Gruschwitz, Lenz & Boehme 1991 [as ephippiata] Haakansson 1981 [as ephippiata] Joger 1981 [as ephippiata] Pauwels & Meirte 1996 [as ephippiata] Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides armitagei Gambian Cylindrical Skink Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Greenbaum 2005 Greenbaum, Campbell & Raxworthy 2006 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Pasteur 1981 Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis tristaoi Nimba Snake-eyed Skink Barnett, Emms & Santoni 2001 [as Leptosiaphis nimbaense] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as nimbense] Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Pauwels & Meirte 1996 Trape, Trape & Chirio 2012   Trachylepis perrotetii Orange-flanked Skink Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Pauwels & Meirte 1996 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Barnett, Emms & Santoni 2001 Bayless 2002 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Barnett, Emms & Santoni 2001 Bayless 2002 Boehme & Ziegler 1997 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Trape, Trape & Chirio 2012

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Genus Dasypeltis (African Egg Eaters)   Dasypeltis fasciata Central African Egg Eater Barnett, Emms & Santoni 2001 Chirio & LeBreton 2007 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gans 1959 Gruschwitz, Lenz & Boehme 1991 Hughes & Barry 1969 Haakansson 1981 Loveridge 1957   Dasypeltis gansi Savanna Egg Eater Chirio & LeBreton 2007 Genus Dispholidus (Boomslangs) Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Pauwels & Meirte 1996   Grayia tholloni Nandi Water Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Pauwels & Meirte 1996 Trape & Mane 2006 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Genus Philothamnus (African Green Snakes)   Philothamnus irregularis Irregular Green Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981   Philothamnus semivariegatus Variegated Bush Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Loveridge 1957 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus variegatus Variable Cat Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis viridis Western Green Mamba Barnett, Emms & Santoni 2001 David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gruschwitz, Lenz & Boehme 1991 Hughes & Barry 1969 Haakansson 1981 Spawls & Branch 1995 Trape & Mane 2006 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Barnett, Emms & Santoni 2001 David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gruschwitz, Lenz & Boehme 1991 Pauwels & Meirte 1996 Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja nigricollis Black-necked Spitting Cobra Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Naja savannula West African Banded Cobra Barnett, Emms & Santoni 2001 [as N. melanoleuca] David & Ineich 1999 [as N. melanoleuca] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as N. melanoleuca] Gruschwitz, Lenz & Boehme 1991 [as N. melanoleuca] Haakansson 1981 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Pauwels & Meirte 1996 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake Chippaux & Jackson 2019 David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Hughes & Barry 1969 Pauwels & Meirte 1996 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis boulengeri Central African Stiletto Snake Laurent 1950   Atractaspis dahomeyensis Dahomey Stiletto Snake David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Pauwels & Meirte 1996   Atractaspis microlepidota Small-scaled Stiletto Snake Chippaux & Jackson 2019 Trape & Mane 2006 Trape 2005 Wallach, Williams & Boundy 2014   Atractaspis micropholis Sahelian Stiletto Snake David & Ineich 1999 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981   Boaedon lineatus Lined House Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Hughes 2004 Genus Lycophidion (African Wolf Snakes)   Lycophidion albomaculatum White-spotted Wolf Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007   Lycophidion irroratum Western Forest Wolf Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Trape & Mane 2006   Lycophidion semicinctum Semi-annulated Wolf Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as L. semicinctum] Gruschwitz, Lenz & Boehme 1991 [as L. semicinctum] Haakansson 1981 [as L. semicinctum] Genus Prosymna (Shovel-snout Snakes)   Prosymna greigerti Sudan Savanna Shovel-snout Chirio & LeBreton 2007 Chirio, Ineich, Schmitz & Trape 2011 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007   Prosymna meleagris Speckled Shovel-snout Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Barnett, Emms & Santoni 2001 [as P. sibilans] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as rukwae] Gruschwitz, Lenz & Boehme 1991 [as rukwae] Haakansson 1981 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Joger 1981   Psammophis phillipsii West African Olive Grass Snake Barnett, Emms & Santoni 2001 Chippaux & Jackson 2019 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis boueti Mali Thread Snake Ineich & Prudent 2014   * Myriopholis natatrix Gambian Thread Snake Andersson 1937 Chippaux & Jackson 2019 Hahn 1980 McDiarmid, Campbell & Toure 1999 Wagner & Wilms 2013 Genus Rhinoleptus (West African Sharp-nosed Thread Snake)   Rhinoleptus koniagui West African Sharp-nosed Thread Snake Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Jallow & Trape 2013  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981   Python sebae Northern Rock Python Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops punctatus Spotted Blind Snake Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Hahn 1980 Haakansson 1981 Wallach, Williams & Boundy 2014  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Barnett, Emms & Santoni 2001 David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus maculatus West African Night Adder David & Ineich 1999 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Pauwels & Meirte 1996 Spawls & Branch 1995 Genus Echis (Carpet Vipers)   Echis jogeri Mali Carpet Viper David & Ineich 1999 [as ocellatus] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 [as ocellatus] Spawls & Branch 1995 [as ocellatus]

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Brongersma in Bjorndal (ed.) 1982 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Pauwels & Meirte 1996 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boycott & Bourquin 2008 [as subrufa] Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 McCord, Joseph-Ouni & Bour 2004 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977   Pelusios castaneus Chestnut Mud Turtle Barnett, Emms & Santoni 2001 Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys nogueyi Western Savanna Hinge-back Tortoise Barnett, Emms & Santoni 2001 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 [as belliana] Haakansson 1981 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis senegalensis Senegal Flapshell Turtle Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Gruschwitz, Lenz & Boehme 1991 Haakansson 1981 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Bonin, Devaux & Dupre 2006 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Haakansson 1981 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012