Reptiles of Ivory Coast    Bibliography of the reptiles of Ivory Coast Taxonomic history of the reptiles of Ivory Coast Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   * Cynisca ivoirensis Gaboua Worm Lizard Mane & Trape 2015 Trape & Mane 2014   Cynisca leucura White-tailed Worm Lizard Gans 1987 Gans 2005   * Cynisca rouxae Assacro Worm Lizard Gans 1987 Gans 2005

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile King & Burke 1989 [as niloticus] Roedel & Grabow 1995 Trape, Trape & Chirio 2012 Genus Mecistops (African Slender-snouted Crocodiles)   Mecistops cataphractus West African Slender-snouted Crocodile Branch & Roedel 2003 King & Burke 1989 Roedel & Branch 2005 Roedel & Grabow 1995 Shirley, Carr, Nestler, Vliet & Brochu 2018 Trape, Trape & Chirio 2012 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Branch & Roedel 2003 King & Burke 1989 Roedel & Branch 2005 Roedel & Grabow 1995 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama picticauda Ada Foah Agama Branch & Roedel 2003 [as agama] Joger 1981 [as agama] Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Roedel & Branch 2005 [as agama] Roedel, Grabow, Hallermann & Boeckheler 1997 [as agama] Trape, Trape & Chirio 2012 [as agama] Wermuth 1967 [as agama]   Agama sankaranica Sankaranian Agama Mediannikov, Trape & Trape 2012 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo gracilis Graceful Chameleon Branch & Roedel 2003 Glaw 2015 Klaver & Boehme 1997 Roedel & Branch 2005 Roedel, Grabow, Hallermann & Boeckheler 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Roedel, Grabow, Hallermann & Boeckheler 1997 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   Ancylodactylus occidentalis West African Rock Gecko Penner & Roedel 2013 Perret 1986 Trape, Trape & Chirio 2012   Ancylodactylus spinicollis Spiny-necked Rock Gecko Perret 1986 Roedel, Grabow, Hallermann & Boeckheler 1997 Segniagbeto 2021 Trape, Trape & Chirio 2012 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus albivertebralis White-lined Half-toed Gecko Boehme & Grell 2013   Hemidactylus angulatus West African House Gecko Ineich 2003 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Hemidactylus ansorgii Slender Forest Gecko Henle & Boehme 2003 Trape, Trape & Chirio 2012   Hemidactylus fasciatus African Banded Forest Gecko Branch & Roedel 2003 Loveridge 1947 Roedel & Branch 2005 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012 Wagner, Leache & Fujita 2014   Hemidactylus mabouia Cosmopolitan House Gecko Trape, Trape & Chirio 2012   Hemidactylus muriceus Prickly Half-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Branch & Roedel 2003 Henle & Boehme 2003 Roedel & Branch 2005 Trape, Trape & Chirio 2012   Hemidactylus pseudomuriceus Azagny Half-toed Gecko Henle & Boehme 2003 Trape, Trape & Chirio 2012 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus conraui Cameroon Dwarf Gecko Barbault 1975 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Gastropholis (Keel-bellied Lizards)   Gastropholis echinata African Spiny-tailed Lizard Arnold 1989 Roedel 1996 Trape, Trape & Chirio 2012 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus togoensis Sahelian Fan-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola ephippiata African Wall Gecko Joger 1984 Roedel, Grabow, Hallermann & Boeckheler 1997 Schleich, Kastle & Kabisch 1996 Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Cophoscincopus (West African Keeled Water Skinks)   Cophoscincopus durus Liberian Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Branch & Roedel 2003 Roedel & Branch 2005 Trape, Trape & Chirio 2012   Cophoscincopus greeri Guinean Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Trape, Trape & Chirio 2012   Cophoscincopus simulans Ghanese Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Trape, Trape & Chirio 2012 Genus Mochlus (African Supple Skinks)   Mochlus brevicaudis Short-tailed Supple Skink Greer, Grandison & Barbault 1985 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Mochlus fernandi Western Red-flanked Skink Trape, Trape & Chirio 2012 Wagner, Boehme, Pauwels & Schmitz 2009   Mochlus guineensis Guinean Supple Skink Grandison 1956 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis togoensis Togo Snake-eyed Skink Perret 1973 Roedel, Grabow, Hallermann & Boeckheler 1997 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012   Panaspis tristaoi Nimba Snake-eyed Skink Barbault 1976 [as nimbense] Fuhn 1972 [as nimbense] Ineich 2003 [as nimbense] Perret 1973 [as nimbense] Roedel, Grabow, Hallermann & Boeckheler 1997 [as nimbense] Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Branch & Roedel 2003 Hoogmoed 1974 Roedel & Branch 2005 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Trachylepis aureogularis Orange-throated Skink Hoogmoed 1974 [as albilabris] Trape, Trape & Chirio 2012   Trachylepis buettneri Togo Savanna Skink Barbault 1976 Hoogmoed 1974 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Trachylepis maculilabris Common Speckled-lipped Skink Barbault 1976 Hoogmoed 1974 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Trachylepis paucisquamis Western Forest Skink Branch & Roedel 2003 Roedel & Branch 2005 Trape, Trape & Chirio 2012   Trachylepis perrotetii Orange-flanked Skink Grandison 1956 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Koppetsch 2020 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Boehme & Ziegler 1997 Roedel, Grabow, Hallermann & Boeckheler 1997 Trape, Trape & Chirio 2012   Varanus stellatus African Forest Monitor Branch & Roedel 2003 [as V. ornatus] Roedel & Branch 2005 [as V. ornatus] Trape, Trape & Chirio 2012 [as V. ornatus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Chippaux & Jackson 2019 Doucet 1963 Trape & Balde 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Doucet 1963 Ineich 2003 Roedel & Mahsberg 2000 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hippocrepis West African Herald Snake Chirio & LeBreton 2007 Rasmussen, Chirio & Ineich 2000 Trape & Balde 2014   Crotaphopeltis hotamboeia Common Herald Snake Doucet 1963 Roedel & Mahsberg 2000 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Chirio & LeBreton 2007 Trape & Balde 2014 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Chirio & LeBreton 2007 Doucet 1963 Gans 1959 Roedel & Mahsberg 2000 Trape & Balde 2014   Dasypeltis gansi Savanna Egg Eater Trape & Balde 2014 Trape & Mane 2006   Dasypeltis parascabra Guinean Egg Eater Trape & Balde 2014 Trape, Mediannikov & Trape 2012 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa guineensis Guinean Nocturnal Tree Snake Rasmussen 1989 [as brevirostris] Roedel & Mahsberg 2000 [as brevirostris] Trape & Balde 2014   Dipsadoboa underwoodi Cameroon Nocturnal Tree Snake Chirio & LeBreton 2007 Rasmussen 1993 Trape & Balde 2014   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Doucet 1963 Ineich 2003 Rasmussen 1993 Roedel & Mahsberg 2000 Trape & Balde 2014   Dipsadoboa viridis Green Nocturnal Tree Snake Doucet 1963 [as elongata] Rasmussen 1993 Trape & Balde 2014 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) David & Ineich 1999 [as D. typus] Doucet 1963 [as D. typus] Roedel, Kouadio & Mahsberg 1999 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Balde 2014 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Branch & Roedel 2003 Doucet 1963 Joger 1981 Roedel & Branch 2005 Roedel & Mahsberg 2000 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Branch & Roedel 2003 Doucet 1963 Jesus, Nagy, Branch, Wink, Brehm & Harris 2009 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014   Hapsidophrys smaragdinus Common Emerald Snake Doucet 1963 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Schatti 1985 Trape & Balde 2014   Meizodon regularis East African Crowned Smooth Snake Schatti 1985 Trape & Balde 2014 Genus Natriciteres (African Marsh Snakes)   Natriciteres fuliginoides Collared Marsh Snake Hughes & Barry 1969   Natriciteres olivacea Olive Marsh Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014   Natriciteres variegata Variable Marsh Snake Branch & Roedel 2003 Doucet 1963 Ineich 2003 Roedel & Branch 2005 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Philothamnus (African Green Snakes)   Philothamnus carinatus Barred Green Snake Doucet 1963 Trape & Balde 2014   Philothamnus heterodermus Emerald Green Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape, Hughes, Collet & Mediannikov 2021   Philothamnus heterolepidotus Slender Green Snake Chirio & LeBreton 2007 Trape & Balde 2014   Philothamnus irregularis Irregular Green Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape, Hughes, Collet & Mediannikov 2021   Philothamnus nitidus Cameroon Green Snake Doucet 1963 Hughes & Barry 1969 Roedel & Mahsberg 2000 Trape & Balde 2014   Philothamnus pobeguini Guinean Green Snake Trape & Balde 2014   Philothamnus semivariegatus Variegated Bush Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999   Philothamnus smithii Smith's Bush Snake Trape & Balde 2014 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Branch & Roedel 2003 Doucet 1963 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Trape & Balde 2014 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus variegatus Variable Cat Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Branch & Roedel 2003 Broadley 2001 Chippaux & Jackson 2019 Doucet 1963 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Thrasops (Black Tree Snakes)   Thrasops occidentalis Western Black Tree Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Doucet 1963 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014   Toxicodryas pulverulenta Western Powdered Cat Snake Branch & Roedel 2003 Doucet 1963 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis polylepis Black Mamba Courtois & Chippeaux 1977 David & Ineich 1999 Foehrenbach 1982 Trape & Balde 2014   Dendroaspis viridis Western Green Mamba David & Ineich 1999 Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1998 Chippaux & Jackson 2019 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Genus Naja (Typical Cobras)   Naja guineensis Black Forest Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja katiensis Brown Spitting Cobra David & Ineich 1999 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Naja nigricollis Black-necked Spitting Cobra Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Naja savannula West African Banded Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Spawls & Branch 1995 [as N. haje] Trape & Balde 2014 Genus Pseudohaje (Tree Cobras)   Pseudohaje goldii Central African Tree Cobra Chippaux & Jackson 2019 Trape & Balde 2014   Pseudohaje nigra Black Tree Cobra David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus lineatus Lined Centipede Eater Trape & Balde 2014 Wallach, Williams & Boundy 2014   Aparallactus lunulatus Reticulated Centipede Eater Chippaux & Jackson 2019 Chirio & LeBreton 2007 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Aparallactus modestus Western Forest Centipede Eater Doucet 1963 Ineich 2003 Roedel & Mahsberg 2000 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Aparallactus niger West African Black Centipede Eater Chippaux & Jackson 2019 Doucet 1963 Trape & Balde 2014 Villiers 1975 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake David & Ineich 1999 Doucet 1963 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis corpulenta Fat Stiletto Snake David & Ineich 1999 Doucet 1963 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Laurent 1950 Roedel & Mahsberg 2000 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake Chippaux & Jackson 2019 David & Ineich 1999 Doucet 1963 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape & Mediannikov 2016   Boaedon lineatus Lined House Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014   Boaedon olivaceus Olive House Snake Doucet 1963 Trape & Balde 2014   Boaedon virgatus West African House Snake Doucet 1963 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus lineatus Red-and-Black Striped Snake Branch & Roedel 2003 Chippaux & Jackson 2019 Doucet 1963 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Doucet 1963 Trape & Balde 2014 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Doucet 1963 Hughes 2004 Trape & Balde 2014   Dromophis praeornatus Ornate Olympic Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Chirio & LeBreton 2007 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014   Gonionotophis klingi Western Forest Dwarf File Snake Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Doucet 1963 Roedel & Mahsberg 2000 Trape & Balde 2014 Genus Kladirostratus (Striped Beaked Snakes)   Kladirostratus togoensis Western Striped Beaked Snake Chirio & Ineich 1991 Chirio & LeBreton 2007 Roedel, Kouadio & Mahsberg 1999 [as acutus] Trape & Balde 2014 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Chirio & LeBreton 2007 Trape & Balde 2014   Limaformosa guirali White-speckled Forest File Snake Doucet 1963 Trape & Balde 2014 Genus Lycophidion (African Wolf Snakes)   Lycophidion irroratum Western Forest Wolf Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014   Lycophidion laterale Masked Wolf Snake Broadley 1996 Hughes & Barry 1969 Trape & Balde 2014   Lycophidion nigromaculatum Black-spotted Wolf Snake Trape & Balde 2014   Lycophidion semicinctum Semi-annulated Wolf Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Doucet 1963 Ineich 2003 Roedel & Mahsberg 2000 Trape & Balde 2014   Mehelya stenophthalmus Small-eyed File Snake Chippaux & Jackson 2019 Trape & Balde 2014 Genus Polemon (African Snake Eaters)   Polemon acanthias Five-striped Snake Eater Branch & Roedel 2003 Doucet 1963 Ineich 2003 Roedel & Branch 2005 Roedel & Mahsberg 2000 Trape & Balde 2014   Polemon barthii Guinea Snake Eater Chippaux & Jackson 2019 Hughes 2003 Roedel & Mahsberg 2000 Trape & Balde 2014   Polemon neuwiedi West African Snake Eater Chippaux & Jackson 2019 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape & Mane 2006 Genus Prosymna (Shovel-snout Snakes)   Prosymna meleagris Speckled Shovel-snout Broadley 1980 Chirio, Ineich, Schmitz & Trape 2011 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Doucet 1963 [as P. sibilans] Roedel, Kouadio & Mahsberg 1999 [as "cf. rukwae"] Trape & Balde 2014 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis phillipsii West African Olive Grass Snake Doucet 1963 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chirio & Ineich 1991 Doucet 1963 Trape & Balde 2014  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis albiventer White-bellied Thread Snake Hallermann & Roedel 1995 McDiarmid, Campbell & Toure 1999 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Trape & Mane 2006   Myriopholis narirostris Sahel Thread Snake Doucet 1963 Hahn 1980 McDiarmid, Campbell & Toure 1999 Papenfuss 1969 Trape & Balde 2014 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma bicolor Bicoloured Thread Snake Doucet 1963 [as brevicauda] Hahn 1980 [as Leptotyphlops brevicaudus] McDiarmid, Campbell & Toure 1999 [as brevicaudus] Papenfuss 1969 [as brevicauda] Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Villiers 1965   * Tricheilostoma broadleyi Lamto Thread Snake Chippaux & Jackson 2019 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Wallach & Hahn 1997  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Doucet 1963 McDiarmid, Campbell & Toure 1999 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014   Python sebae Northern Rock Python Branch & Roedel 2003 Broadley 1984 Doucet 1963 Roedel & Branch 2005 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops liberiensis Liberian Blind Snake Branch & Roedel 2003 Chippaux & Jackson 2019 Doucet 1963 [as Typhlops hallowelli] Hahn 1980 Roedel & Branch 2005 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops lineolatus Lineolate Blind Snake Hahn 1980 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Hahn 1980 Roedel, Kouadio & Mahsberg 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Typhlops (Afro-Caribbean Blind Snakes)   Typhlops coecatus Ghanese Blind Snake Chippaux & Jackson 2019 Doucet 1963 Hahn 1980 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   Atheris chlorechis West African Green Bush Viper Branch & Roedel 2003 Chippaux & Jackson 2019 David & Ineich 1999 Doucet 1963 Ineich 2003 McDiarmid, Campbell & Toure 1999 Roedel & Branch 2005 Roedel & Mahsberg 2000 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atheris hirsuta Tai Hairy Bush Viper Chippaux & Jackson 2019 Ernst & Roedel 2002 Penner, Gonwouo & Roedel 2013 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Chippaux & Jackson 2019 David & Ineich 1999 Doucet 1963 McDiarmid, Campbell & Toure 1999 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014   Bitis nasicornis African Nose-horned Viper David & Ineich 1999 Doucet 1963 Roedel & Mahsberg 2000 Spawls & Branch 1995 Trape & Balde 2014   Bitis rhinoceros Rhinoceros Viper Chippaux & Jackson 2019 Doucet 1963 [as gabonica] Roedel & Mahsberg 2000 [as gabonica] Spawls & Branch 1995 [as gabonica] Trape & Balde 2014 Genus Causus (Night Adders)   Causus lichtensteinii Forest Night Adder Chippaux & Jackson 2019 David & Ineich 1999 Roedel & Mahsberg 2000 Spawls & Branch 1995 Trape & Balde 2014   Causus maculatus West African Night Adder Branch & Roedel 2003 David & Ineich 1999 Roedel & Branch 2005 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Genus Echis (Carpet Vipers)   Echis ocellatus West African Carpet Viper David & Ineich 1999 Hughes 1976 McDiarmid, Campbell & Toure 1999 Roedel, Kouadio & Mahsberg 1999 Spawls & Branch 1995 Trape & Balde 2014 Trape 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roedel & Grabow 1995 Trape, Trape & Chirio 2012   Pelomedusa variabilis West African Helmeted Turtle Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Chirio & LeBreton 2007 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roedel & Grabow 1995 Trape, Trape & Chirio 2012   Pelusios cupulatta Ivorian Mud Turtle Bour & Maran 2003 Maran 2009 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios gabonensis Forest Mud Turtle Chirio & LeBreton 2007 Trape, Trape & Chirio 2012  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys erosa Forest Hinge-back Tortoise Branch & Roedel 2003 King & Burke 1989 Luiselli & Diagne 2014 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roedel & Branch 2005 Trape, Trape & Chirio 2012   Kinixys homeana West African Hinge-back Tortoise King & Burke 1989 Luiselli & Diagne 2013 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys nogueyi Western Savanna Hinge-back Tortoise Maran 2009 McCord, Joseph-Ouni & Tabaka 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roedel & Grabow 1995 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis senegalensis Senegal Flapshell Turtle Chirio & LeBreton 2007 King & Burke 1989 Maran 2009 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roedel & Grabow 1995 Trape, Trape & Chirio 2012 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle King & Burke 1989 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012