Reptiles of Senegal    Bibliography of the reptiles of Senegal Taxonomic history of the reptiles of Senegal Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca feae Guinea-Bissau Worm Lizard Gans 1987 Gans 2005 Mane & Trape 2015 Trape 2014   Cynisca manei Bamako Samba Worm Lizard Mane & Trape 2015 Trape 2014 Trape, Mane & Balde 2014   Cynisca senegalensis Senegal Worm Lizard Gans 1987 Gans 2005 Mane & Trape 2015 Trape 2014

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Brito 2003 [as niloticus] Fuchs, Mertens & Wermuth 1974 Joger & Lambert 2002 [as niloticus] King & Burke 1989 [as niloticus] Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Joger & Lambert 2002 Spawls, Howell, Drewes & Ashe 2002 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama bocourti Casamance Agama The Reptile Database (viewed 10th April, 2011) Wermuth 1967   Agama boensis Boe Agama Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012   Agama boueti Mali Agama Berre 1989 Boehme 1978 Cisse & Karns 1978 Joger 1979 Trape, Trape & Chirio 2012   Agama picticauda Ada Foah Agama Cisse & Karns 1978 [as agama] Geniez, Padial & Crochet 2011 [as agama] Joger & Lambert 2002 [as agama] Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Mediannikov, Trape & Trape 2012 [as agama] Monasterio, Alvarez, Trape & Roedel 2016 [as agama] Trape, Trape & Chirio 2012 [as agama] Wagner, Krause & Boehme 2008 [as agama]   Agama sankaranica Sankaranian Agama Joger & Lambert 2002   Agama weidholzi Senegal Agama Cisse & Karns 1978 Joger & Lambert 1996 Joger & Lambert 2002 Joger 1979 Joger 1981 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Wagner & Bauer 2011 Wagner, Ineich, Leache, Wilms, Trape, Boehme & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo gracilis Graceful Chameleon Boehme 1978 Cisse & Karns 1978 Glaw 2015 Joger & Lambert 2002 Klaver & Boehme 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Cisse & Karns 1978 Glaw 2015 Joger & Lambert 2002 Joger 1981 Klaver & Boehme 1997 Monasterio, Alvarez, Trape & Roedel 2016 Papenfuss 1969 Tilbury 2010 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Cisse & Karns 1978 Grismer 1988 Joger & Lambert 2002 Joger 1981 Loveridge 1947 Papenfuss 1969 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Boehme 1978 [as brookii] Cisse & Karns 1978 [as brookii] Joger & Lambert 2002 [as "brooki complex"] Loveridge 1947 Monasterio, Alvarez, Trape & Roedel 2016 Trape, Trape & Chirio 2012 Wermuth 1965   Hemidactylus mabouia Cosmopolitan House Gecko Boehme 1978 Cisse & Karns 1978 Trape, Trape & Chirio 2012 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Boehme 1978 Cisse & Karns 1978 Joger & Lambert 1996 Trape, Trape & Chirio 2012 Wermuth 1965 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus petrii North African Thin-toed Gecko Arnold 1980 Cisse & Karns 1978 Trape, Trape & Chirio 2012   Stenodactylus sthenodactylus Elegant Thin-toed Gecko Baha el Din 2006 Boehme 1978 Sindaco & Jeremchenko 2008  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus aureus Golden Fringe-toed Lizard Crochet, Geniez & Ineich 2003 Salvador 1982 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Acanthodactylus dumerilii Lesser Fringe-toed Lizard Crochet, Geniez & Ineich 2003 [as senegalensis] Salvador 1982 Schleich, Kastle & Kabisch 1996 Trape, Trape & Chirio 2012 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Bischoff 1991 Boehme 1978 Cisse & Karns 1978 Papenfuss 1969 Trape, Trape & Chirio 2012 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina guttulata Common Small-spotted Desert Lizard Schleich, Kastle & Kabisch 1996 Szczerbak 1975 Yousefkhani, Marin & Rastegar-Pouyani 2015  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Cisse & Karns 1978 Joger 1981 Joger 1984 Papenfuss 1969 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola chazaliae Helmeted Gecko Bons & Geniez 1996 Geniez, Mateo, Geniez & Pether 2004 Ineich, Dia, Alphally & Colas 1998   Tarentola parvicarinata Small-keeled Gecko Berre 1989 Joger & Lambert 1996 Joger & Lambert 2002 Joger 1984 Trape, Trape & Chirio 2012   Tarentola senegambiae Senegambian Wall Gecko Cisse & Karns 1978 [as ephippiata] Joger & Lambert 2002 [as ephippiata] Joger 1981 [as ephippiata] Joger 1984 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides armitagei Gambian Cylindrical Skink Carranza, Arnold, Geniez, Roca & Mateo 2008 Trape 2008 Trape, Trape & Chirio 2012   Chalcides delislei Central Saharan Three-fingered Skink Papenfuss 1969 Trape, Trape & Chirio 2012   Chalcides pulchellus West African Cylindrical Skink Carranza, Arnold, Geniez, Roca & Mateo 2008 Greenbaum 2005 Joger & Lambert 2002 Trape, Trape & Chirio 2012   Chalcides sphenopsiformis Senegal Sand Skink Boehme 1978 Cisse & Karns 1978 Grandison 1956 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Cophoscincopus (West African Keeled Water Skinks)   Cophoscincopus greeri Guinean Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Joger & Lambert 2002 [as "cf. greeri"]   Cophoscincopus senegalensis Senegalese Keeled Water Skink Trape, Trape & Chirio 2012 Genus Mochlus (African Supple Skinks)   Mochlus fernandi Western Red-flanked Skink Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis togoensis Togo Snake-eyed Skink Boehme 1978 Fuhn 1972 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012   Panaspis tristaoi Nimba Snake-eyed Skink Trape 2010 [as P. "nimbaense"] Trape, Trape & Chirio 2012 Genus Scincus (Sand Fish)   Scincus albifasciatus White-banded Sand Fish Arnold & Leviton 1977 Boehme 1978 Cisse & Karns 1978 Geniez, Mateo, Geniez & Pether 2004 Hartmann 1989 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Cisse & Karns 1978 Hoogmoed 1974 Joger 1981 Monasterio, Alvarez, Trape & Roedel 2016 Trape, Trape & Chirio 2012   Trachylepis perrotetii Orange-flanked Skink Cisse & Karns 1978 Hoogmoed 1974 Joger & Lambert 2002 Joger 1981 Papenfuss 1969 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Hoogmoed 1974 Joger & Lambert 2002 Koppetsch 2020 Papenfuss 1969 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Boehme 1978 Cisse & Karns 1978 Mertens 1963 Papenfuss 1969 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Boehme & Ziegler 1997 Joger & Lambert 2002 Monasterio, Alvarez, Trape & Roedel 2016 Trape, Trape & Chirio 2012

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Berre 1989 Boehme 1978 Stimson 1969 Trape & Mane 2002 Trape & Mane 2006  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Chippaux & Jackson 2019 Joger & Lambert 2002 Monasterio, Alvarez, Trape & Roedel 2016 Trape & Mane 1995 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Wallach, Williams & Boundy 2014 Genus Bamanophis (West African Whip Snake)   Bamanophis dorri West African Whip Snake Chippaux & Jackson 2019 Joger & Lambert 2002 Papenfuss 1969 Schatti & Trape 2008 Trape & Mane 2002 Trape & Mane 2004 Trape 1997 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Boehme 1978 Joger & Lambert 2002 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Bates & Broadley 2018 Chippaux & Jackson 2019 Trape & Mane 2006 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Chirio & LeBreton 2007 Gans 1959 Gans 1960 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004   Dasypeltis gansi Savanna Egg Eater Bates & Ineich 2012 Bates, Trape & Spawls 2013 Chippaux & Jackson 2019 Trape & Mane 2006 Trape, Mediannikov & Trape 2012   Dasypeltis latericia Senegalese Egg Eater Chippaux & Jackson 2019 Trape & Mane 2006 Trape, Mediannikov & Trape 2012   Dasypeltis sahelensis Sahel Egg Eater Chippaux & Jackson 2019 Sindaco, Venchi & Grieco 2013 Trape & Mane 2006 Trape, Mediannikov & Trape 2012 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa weileri Grey Nocturnal Tree Snake Monasterio, Alvarez, Trape & Roedel 2016 [as "aff. weileri"] Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) Broadley & Wallach 2002 [as D. typus] David & Ineich 1999 [as D. typus] Joger & Lambert 2002 [as D. t. typus] Papenfuss 1969 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Mane 2002 [as D. typus] Trape & Mane 2006 [as D. typus] Villiers 1975 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Chippaux & Jackson 2019 Joger & Lambert 1996 Joger & Lambert 2002 Joger 1981 Monasterio, Alvarez, Trape & Roedel 2016 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006   Grayia tholloni Nandi Water Snake Chippaux & Jackson 2019 Pauwels 1996 Trape & Mane 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Chippaux & Jackson 2019 Joger & Lambert 2002 Papenfuss 1969 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975   Meizodon semiornatus Semi-ornate Smooth Snake Doucet 1963 Villiers 1975 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Trape & Mane 2006 Genus Philothamnus (African Green Snakes)   Philothamnus irregularis Irregular Green Snake Boehme 1978 Chippaux & Jackson 2019 Joger & Lambert 2002 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006   Philothamnus semivariegatus Variegated Bush Snake Boehme 1978 Chippaux & Jackson 2019 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006   Philothamnus smithii Smith's Green Snake Trape, Hughes, Collet & Mediannikov 2021 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus tripolitanus North African Cat Snake Chippaux & Jackson 2019 Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Papenfuss 1969 Sindaco, Venchi & Grieco 2013 Trape & Mane 2002 Trape & Mane 2006 Wilms, Wagner & Spawls 2021   Telescopus variegatus Variable Cat Snake Chippaux & Jackson 2019 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Genus Thrasops (Black Tree Snakes)   Thrasops occidentalis Western Black Tree Snake Chippaux & Jackson 2019 Trape & Mane 2002 Trape & Mane 2006 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Doucet 1963 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Trape & Mane 2002 Trape & Mane 2006 Villiers 1975  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis polylepis Black Mamba David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haakansson & Madsen 1983 Spawls & Branch 1995 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Trape, Mane & Balde 2005 Villiers 1975   Dendroaspis viridis Western Green Mamba Chippaux & Jackson 2019 David & Ineich 1999 Trape & Mane 2002 Trape & Mane 2006 Villiers 1975 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1971 Broadley 1998 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006   Elapsoidea trapei Senegal Garter Snake Chippaux & Jackson 2019 Mane 1999 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Genus Naja (Typical Cobras)   Naja katiensis Brown Spitting Cobra Chippaux & Jackson 2019 David & Ineich 1999 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Wallach, Williams & Boundy 2014   Naja nigricollis Black-necked Spitting Cobra Broadley 1968 Joger & Lambert 2002 Monasterio, Alvarez, Trape & Roedel 2016 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Wallach, Williams & Boundy 2014 Wuster & Broadley 2007   Naja savannula West African Banded Cobra Boehme 1978 [as N. melanoleuca] Broadley 1968 [as N. melanoleuca] David & Ineich 1999 [as N. melanoleuca] Monasterio, Alvarez, Trape & Roedel 2016 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Trape & Mane 2000 [as N. melanoleuca] Trape & Mane 2002 [as N. melanoleuca] Trape & Mane 2004 [as N. melanoleuca] Trape & Mane 2006 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Broadley 1968 [as N. haje] Chippaux & Jackson 2019 Spawls & Branch 1995 [as N. haje] Trape & Mane 2000 [as N. haje] Trape & Mane 2002 [as N. haje] Trape & Mane 2004 [as N. haje] Trape & Mane 2006 [as N. haje] Trape, Chirio, Broadley & Wuster 2009  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Chippaux & Jackson 2019 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake Boehme 1978 David & Ineich 1999 Spawls & Branch 1995 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake Chippaux & Jackson 2019 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Joger & Lambert 2002 Spawls & Branch 1995 Villiers 1975 Wallach, Williams & Boundy 2014   Atractaspis microlepidota Small-scaled Stiletto Snake David & Ineich 1999 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006 Trape 2005 Trape, Mane & Ineich 2006 Wallach, Williams & Boundy 2014   Atractaspis micropholis Sahelian Stiletto Snake Chippaux & Jackson 2019 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006 Trape, Mane & Ineich 2006 Villiers 1975 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Joger & Lambert 2002 Joger 1981 Papenfuss 1969 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975   Boaedon lineatus Lined House Snake Chippaux & Jackson 2019 Joger & Lambert 2002 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Chippaux & Jackson 2019 Hughes 2004 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006 Villiers 1975   Dromophis praeornatus Ornate Olympic Snake Boehme 1978 Chippaux & Jackson 2019 Monasterio, Alvarez, Trape & Roedel 2016 Papenfuss 1969 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Villiers 1965 Villiers 1975 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Chippaux & Jackson 2019 Trape & Mane 2002 Trape & Mane 2006 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Chippaux & Jackson 2019 Papenfuss 1969 Trape & Mane 2002 Trape & Mane 2006 Villiers 1965 Genus Lycophidion (African Wolf Snakes)   Lycophidion albomaculatum White-spotted Wolf Snake Chippaux & Jackson 2019 Condamin 1994 Joger & Lambert 1996 Joger & Lambert 2002 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006   Lycophidion irroratum Western Forest Wolf Snake Papenfuss 1969 Villiers 1975   Lycophidion semicinctum Semi-annulated Wolf Snake Condamin 1994 [as L. semicinctum] Papenfuss 1969 [as L. semicinctum] Trape & Mane 2002 [as L. semicinctum] Villiers 1975 [as L. semicinctum] Genus Mehelya (Greater File Snakes)   * Mehelya gabouensis Gabou File Snake Chippaux & Jackson 2019 Trape & Mane 2005 Trape & Mane 2006 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chirio, Ineich, Schmitz & Trape 2011 Trape & Mane 2006   Prosymna greigerti Sudan Savanna Shovel-snout Chippaux & Jackson 2019 Chirio, Ineich, Schmitz & Trape 2011 Joger & Lambert 2002 Papenfuss 1969 [as P. laurenti] Trape & Mane 2002 Trape & Mane 2006   Prosymna meleagris Speckled Shovel-snout Chirio, Ineich, Schmitz & Trape 2011 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006 Villiers 1975 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Boehme 1978 [as P. sibilans] Brito, Rebelo, Crochet & Geniez 2008 [as P. sibilans] Joger 1981 [as "cf. rukwae"] Trape & Mane 2000 [as P. sibilans] Trape & Mane 2002 [as P. sibilans] Trape & Mane 2004 [as P. sibilans] Trape & Mane 2006 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Joger & Lambert 2002 Joger 1981 Monasterio, Alvarez, Trape & Roedel 2016 Papenfuss 1969 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975   Psammophis phillipsii West African Olive Grass Snake Broadley 1977 Joger & Lambert 2002 Joger 1981 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis sudanensis Sudan Sand Snake Chippaux & Jackson 2019 Emms, Jambang, Bahl, Mankali, Paziaud & Barnett 2007 Trape & Mane 2002 Trape & Mane 2006 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chippaux & Jackson 2019 Chirio & Ineich 1991 Joger & Lambert 2002 Joger 1981 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis adleri Chadian Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Trape 2002   Myriopholis boueti Mali Thread Snake Hahn & Wallach 1998 Hahn 1980 Ineich & Prudent 2014 Papenfuss 1969 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2006   Myriopholis rouxestevae Senegal Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Trape & Mane 2004 Trape & Mane 2006 Genus Rhinoleptus (West African Sharp-nosed Thread Snake)   Rhinoleptus koniagui West African Sharp-nosed Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Hahn 1980 Jallow & Trape 2013 McDiarmid, Campbell & Toure 1999 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Trape 2014  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Boehme 1978 Chippaux & Jackson 2019 McDiarmid, Campbell & Toure 1999 Monasterio, Alvarez, Trape & Roedel 2016 Papenfuss 1969 Stimson 1969 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Villiers 1975   Python sebae Northern Rock Python Boehme 1978 Broadley 1984 Chippaux & Jackson 2019 Joger & Lambert 2002 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Villiers 1975  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops lineolatus Lineolate Blind Snake Hahn 1980 Joger & Lambert 2002 McDiarmid, Campbell & Toure 1999 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Hahn 1980 Joger & Lambert 2002 Papenfuss 1969 Trape & Mane 2002 Trape & Mane 2006 Wallach, Williams & Boundy 2014  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Boehme 1978 David & Ineich 1999 Joger & Lambert 2002 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Genus Causus (Night Adders)   Causus maculatus West African Night Adder David & Ineich 1999 Joger & Lambert 2002 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002 Trape & Mane 2004 Trape & Mane 2006 Genus Echis (Carpet Vipers)   Echis jogeri Mali Carpet Viper Boehme 1978 [as ocellatus] Chippaux & Jackson 2019 David & Ineich 1999 [as ocellatus] Hughes 1976 [as ocellatus] McDiarmid, Campbell & Toure 1999 [as ocellatus] Monasterio, Alvarez, Trape & Roedel 2016 Pook, Joger, Stumpel & Wuster 2009 Spawls & Branch 1995 [as ocellatus] Trape & Mane 2002 [as ocellatus] Trape & Mane 2004 [as ocellatus] Trape & Mane 2006 [as ocellatus] Trape & Mane 2017 Trape 2018   Echis leucogaster White-bellied Carpet Viper Boehme 1978 David & Ineich 1999 Hughes 1976 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Trape & Mane 2000 Trape & Mane 2002

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Pritchard 1979 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Brongersma in Bjorndal (ed.) 1982 Pritchard 1979 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Brongersma in Bjorndal (ed.) 1982 Fretey 1991 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boehme 1978 [as subrufa] Boycott & Bourquin 2008 [as subrufa] Fritz & Havas 2006 [as subrufa] Iverson 1992 [as subrufa] Joger & Lambert 2002 [as subrufa] King & Burke 1989 [as subrufa] McCord, Joseph-Ouni & Bour 2004 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Boehme 1978 Bour 2008 Fritz & Havas 2006 Iverson 1992 King & Burke 1989 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Fritz & Havas 2006 Iverson 1992 King & Burke 1989 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios castanoides Yellow-bellied Mud Turtle King & Burke 1989   Pelusios subniger East African Black Mud Turtle Joger & Lambert 2002 Joger 1981 King & Burke 1989  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Papenfuss 1969 Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Kinixys (Hinge-back Tortoises)   Kinixys nogueyi Western Savanna Hinge-back Tortoise Fritz & Havas 2006 Iverson 1992 Joger & Lambert 2002 Kindler, Branch, Hofmeyr, Maran, Siroky, Vences, Harvey, Hauswaldt, Schleicher, Stuckas & Fritz 2012 King & Burke 1989 [as belliana] McCord, Joseph-Ouni & Tabaka 2005 Monasterio, Alvarez, Trape & Roedel 2016 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 [as belliana] Trape, Trape & Chirio 2012 [as belliana] Wermuth & Mertens 1977  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis senegalensis Senegal Flapshell Turtle Fritz & Havas 2006 Iverson 1992 Joger & Lambert 2002 King & Burke 1989 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Papenfuss 1969 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Papenfuss 1969 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012