Reptiles of Sudan    Bibliography of the reptiles of Sudan Taxonomic history of the reptiles of Sudan Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Fuchs, Mertens & Wermuth 1974 Wermuth & Mertens 1977 Wilson 1978

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama doriae Nigerian Agama Moody & Boehme 1984   Agama hartmanni Dongola Agama Wagner & Bauer 2011 Werner 1908   Agama picticauda Ada Foah Agama Werner 1908 [as colonorum]   Agama rueppelli Arboreal Agama Wagner & Bauer 2011   Agama spinosa Egyptian Agama Barbour 1913 Berre 1989 [as agama] Geniez, Padial & Crochet 2011 Wagner, Leache, Mazuch & Boehme 2013 Werner 1908 Genus Pseudotrapelus (Afro-Arabian Desert Agamas)   Pseudotrapelus chlodnickii North East African Desert Agama Berre 1989 [as sinaitus] Melnikov, Smielowski, Melnikova, Nazarov & Ananjeva 2015 Tamar, Scholz, Crochet, Geniez, Meiri, Schmitz, Wilms & Carranza 2016 Werner 1908 [as sinaitus] Genus Trapelus (Afro-Asian Ground Agamas)   Trapelus mutabilis North African Desert Agama Berre 1989 Capocaccia 1977 Werner 1908 [as Agama pallida] Genus Uromastyx (Western Spiny-tailed Agamas)   Uromastyx dispar South Saharan Spiny-tailed Agama Mohammed & Hammad 2008 Wermuth 1967 Werner 1908 [as acanthinurus] Wilms & Boehme 2001 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   Uromastyx ocellata Ocellated Spiny-tailed Agama Berre 1989 Lanza 1990 Largen 1997 Mohammed & Hammad 2008 Werner 1908 Wilms & Boehme 2000 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo africanus Sahel Chameleon Barbour 1913 [as basiliscus] Berre 1989 Glaw 2015 Werner 1908 [as basiliscus]   Chamaeleo laevigatus Smooth Chameleon Glaw 2015 Werner 1908  Family Gerrhosauridae (Plated Lizards) Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Loveridge 1942 Werner 1908  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Berre 1989 Salvador 1982 Werner 1908   Acanthodactylus scutellatus Nidua Fringe-toed Lizard Arnold 1983 Berre 1989 Crochet, Geniez & Ineich 2003 Salvador 1982 Werner 1908 Genus Heliobolus (Bushveld Lizards)   Heliobolus spekii East African Bushveld Lizard Szczerbak 1975 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Barbour 1913 Berre 1989 Werner 1908 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina guttulata Common Small-spotted Desert Lizard Abukashawa & Mukhtar 2015 Berre 1989 Sindaco, Simo-Riudalbas, Sacchi & Carranza 2018 Szczerbak 1975 Werner 1908 Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina martini Red Sea Desert Lizard Largen 1997 Szczerbak 1975 Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina rubropunctata Red-spotted Desert Lizard Berre 1989 Bischoff 1991 Szczerbak 1975 Yousefkhani, Marin & Rastegar-Pouyani 2015 Genus Philochortus (Shield-backed Ground Lizards)   Philochortus intermedius Northern Shield-backed Ground Lizard Berre 1989 Genus Pseuderemias (Sand Runners)   Pseuderemias brenneri Barawa Sand Runner Bischoff 1992 Szczerbak 1975   Pseuderemias mucronata Anseba Sand Runner Berre 1989 Bischoff 1992 Szczerbak 1975 Werner 1908  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus hasselquistii Egyptian Fan-toed Gecko Heimes 1987 Loveridge 1947 Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015 Werner 1908   Ptyodactylus togoensis Sahelian Fan-toed Gecko Heimes 1987 [as P. ragazzii] Schleich, Kastle & Kabisch 1996 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Barbour 1913 Berre 1989 Joger 1984 Werner 1908   Tarentola ephippiata African Wall Gecko Berre 1989 Joger 1984 Schleich, Kastle & Kabisch 1996 Werner 1908  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides bottegi Ethiopian Cylindrical Skink Greenbaum, Campbell & Raxworthy 2006   Chalcides delislei Central Saharan Three-fingered Skink Berre 1989 Carranza, Arnold, Geniez, Roca & Mateo 2008 Schleich, Kastle & Kabisch 1996 Werner 1908   Chalcides humilis Saharan Ocellated Skink Carranza, Arnold, Geniez, Roca & Mateo 2008   Chalcides ocellatus Mediterranean Ocellated Skink Barbour 1913 Berre 1989 Greenbaum, Campbell & Raxworthy 2006 Werner 1908 Genus Panaspis (African Snake-eyed Skinks)   * Panaspis wilsoni Sudan Snake-eyed Skink Fuhn 1970 Genus Scincopus (Tiger Skinks)   Scincopus fasciatus Banded Tiger Skink Berre 1989 Lanza & Corsi 1981 Schleich, Kastle & Kabisch 1996 Werner 1908   * Scincopus melanocephalus Black-headed Tiger Skink Lanza & Corsi 1981 Sindaco 1995 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis brevicollis Short-necked Skink Spawls, Howell, Hinkel & Menegon 2018 Werner 1908   Trachylepis quinquetaeniata African Five-lined Skink Barbour 1913 Berre 1989 Koppetsch 2020 Werner 1908   Trachylepis striata Eastern Striped Skink Werner 1908  Family Sphaerodactylidae (Least Geckos, etc.) Genus Pristurus (Semaphore Geckos)   Pristurus flavipunctatus Yellow-spotted Semaphore Gecko Arnold 2009 Berre 1989 Loveridge 1947 Werner 1908  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bayless 2002   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Werner 1908 [as V. ocellatus]   Varanus griseus Desert Monitor Bayless 2002 Werner 1908   Varanus niloticus Nile Monitor Bayless 2002 Berre 1989 Werner 1908

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx colubrinus East African Sand Boa Broadley 1994 Colley 1946 [as E. thebaicus] Corkill 1935 Sindaco, Venchi & Grieco 2013 Werner 1908 [as E. thebaicus]   Eryx muelleri West African Sand Boa Berre 1989 Werner 1908  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis degeni Yellow-flanked Herald Snake Chippaux & Jackson 2019 Werner 1908 [as Leptodira attarensis]   Crotaphopeltis hotamboeia Common Herald Snake Corkill 1935 Genus Dasypeltis (African Egg Eaters)   Dasypeltis sahelensis Sahel Egg Eater Colley 1946 [as D. scabra] Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020   Dasypeltis scabra Common African Egg Eater Bates & Broadley 2018 Corkill 1935 Werner 1908 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) Corkill 1935 [as D. typus] Spawls & Branch 1995 [as D. typus] Genus Eirenis (Middle Eastern Dwarf Snakes)   Eirenis africanus African Dwarf Snake Corkill 1935 Largen & Rasmussen 1993 Largen 1997 Mahlow, Tillack, Schmidtler & Muller 2013 Genus Lytorhynchus (Awl-headed Snakes)   Lytorhynchus diadema Crowned Awl-headed Snake Sindaco, Venchi & Grieco 2013 Werner 1908 Genus Meizodon (African Smooth Snakes)   Meizodon semiornatus Semi-ornate Smooth Snake Corkill 1935 Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Loveridge 1958 Osman & El Sir 1988 Werner 1908 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Largen & Rasmussen 1993 Werner 1908 [as Chlorophis emini]   Philothamnus bequaerti Uganda Green Snake Corkill 1935 [as Chlorophis irregularis] Loveridge 1958 [as i. irregularis]   Philothamnus semivariegatus Variegated Bush Snake Corkill 1935 Loveridge 1958 Pitman 1974 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps florulentus Egyptian Whip Snake Barbour 1913 Broadley 1994 Corkill 1935 Osman & El Sir 1988 Schatti 1988 Sindaco, Venchi & Grieco 2013 Werner 1908   Platyceps saharicus Saharan Cliff Racer Schatti & McCarthy 2004 Sindaco, Venchi & Grieco 2013 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Corkill 1935   Scaphiophis raffreyi Ethiopian Hook-nosed Snake Broadley 1994 Genus Spalerosophis (Diadem Snakes)   Spalerosophis diadema Common Diadem Snake Berre 1989 Corkill 1935 Osman & El Sir 1988 Sindaco, Venchi & Grieco 2013 Werner 1908 Genus Telescopus (Afro-Asian Cat Snakes)   * Telescopus gezirae Blue Nile Cat Snake Broadley 1994 Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008   Telescopus obtusus Egyptian Cat Snake Berre 1989 Chippaux & Jackson 2019 Corkill 1935 [as Tarbophis obtusus] Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Osman & El Sir 1988 Sindaco, Venchi & Grieco 2013 [as dhara] Spawls, Malonza & Beraduccii 2021 Werner 1908   Telescopus tripolitanus North African Cat Snake Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Sindaco, Venchi & Grieco 2013 Wilms, Wagner & Spawls 2021  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis polylepis Black Mamba Spawls & Branch 1995 Genus Elapsoidea (African Garter Snakes)   Elapsoidea laticincta Sudanese Garter Snake Broadley 1971 Chippaux & Jackson 2019 Loveridge 1944 Genus Naja (Typical Cobras)   Naja haje Egyptian Cobra Broadley 1994 Colley 1946 Corkill 1935 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Trape, Chirio, Broadley & Wuster 2009 Werner 1908   Naja nigricollis Black-necked Spitting Cobra Berre 1989 Corkill 1935 Werner 1908 Wuster & Broadley 2003 Wuster & Broadley 2007 [as "cf. nigricollis"]   Naja nubiae Nubian Spitting Cobra Chippaux & Jackson 2019 Sindaco, Venchi & Grieco 2013 Wallach, Williams & Boundy 2014 Wuster & Broadley 2003  Family Lamprophiidae (House Snakes, etc.) Genus Aparallactus (African Centipede Eaters)   Aparallactus lunulatus Reticulated Centipede Eater Pitman 1974 Genus Atractaspis (Stiletto Snakes)   Atractaspis magrettii Mandafena Stiletto Snake Largen & Spawls 2010 Largen 1997 Wallach, Williams & Boundy 2014   Atractaspis phillipsi Sudanese Stiletto Snake Barbour 1913 Broadley 1994 [as A. microlepidota] Colley 1946 [as A. microlepidota] Corkill 1935 [as A. microlepidota] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 [as A. microlepidota] Wallach, Williams & Boundy 2014   Atractaspis watsoni Sokoto Stiletto Snake Chippaux & Jackson 2019 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Corkill 1935   Boaedon lineatus Lined House Snake Colley 1946   Boaedon longilineatus Long-lined House Snake Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020 Genus Limaformosa (Venus File Snakes)   Limaformosa savorgnani Central African Forest File Snake Corkill 1935 [as chanleri] Lanza & Broadley 2014 Genus Lycophidion (African Wolf Snakes)   Lycophidion jacksoni Kilimanjaro Wolf Snake Broadley & Howell 1991 Broadley & Hughes 1993 Corkill 1935 [as L. capense] Laurent 1968 Trape 2021 Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Berre 1989 [as M. monspessulanus] Schleich, Kastle & Kabisch 1996 Wallach, Williams & Boundy 2014   Malpolon moilensis Moila Snake Berre 1989 Colley 1946 Corkill 1935 Osman & El Sir 1988 Sindaco, Venchi & Grieco 2013 Werner 1908 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chirio, Ineich, Schmitz & Trape 2011   Prosymna greigerti Sudan Savanna Shovel-snout Chirio, Ineich, Schmitz & Trape 2011 Hughes 1983 [as P. meleagris] Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis mossambicus East African Olive Grass Snake Broadley 1994 [as phillipsii] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis punctulatus Northern Speckled Sand Snake Corkill 1935 Loveridge 1940 Sindaco, Venchi & Grieco 2013   Psammophis rukwae Lake Rukwa Sand Snake Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis schokari Arabian Sand Snake Berre 1989 Corkill 1935 Loveridge 1940 Osman & El Sir 1988 Werner 1908   Psammophis sibilans Hissing Sand Snake Barbour 1913 Berre 1989 Broadley 1994 Corkill 1935 Loveridge 1940 Osman & El Sir 1988 Sindaco, Venchi & Grieco 2013 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Werner 1908   Psammophis sudanensis Sudan Sand Snake Chippaux & Jackson 2019 Colley 1946 [as P. subtaeniatus] Corkill 1935 [as P. subtaeniatus] Loveridge 1940 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis tanganicus Tanganyika Sand Snake Corkill 1935 [as P. biseriatus] Loveridge 1940 Werner 1908 [as P. biseriatus] Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis rostratus Rufous Beaked Snake Corkill 1935  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis braccianii Eritrean Mountain Thread Snake Broadley & Wallach 2007 Hahn & Wallach 1998 [as cairi] Loveridge 1955 [as cairi] Sindaco, Venchi & Grieco 2013 [as cairi]   Myriopholis cairi Egyptian Thread Snake Broadley & Wallach 2007 Corkill 1935   Myriopholis macrorhyncha Large-snouted Thread Snake Broadley & Wallach 2007 Hahn 1978 Sindaco, Venchi & Grieco 2013 Werner 1908 Genus Tricheilostoma (West African Forest Thread Snakes)   * Tricheilostoma dissimilis Sudan Thread Snake Broadley & Wallach 2007  Family Micrelapidae (Short-jawed Rear-fanged Snakes) Genus Micrelaps (Afro-Asian Black-headed Snakes)   Micrelaps vaillanti Somali Black-headed Snake Rasmussen 2002  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Chippaux & Jackson 2019 Corkill 1935   Python sebae Northern Rock Python Broadley 1984 Corkill 1935 Pitman 1974 Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops lineolatus Lineolate Blind Snake Broadley & Wallach 2009 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Corkill 1935  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Broadley 1994 Corkill 1935 Osman & El Sir 1988 Spawls & Branch 1995 Werner 1908 Genus Causus (Night Adders)   Causus resimus Green Night Adder Broadley 1994 Chippaux & Jackson 2019 Corkill 1935 Pitman 1974 Werner 1908   Causus rhombeatus Rhombic Night Adder Corkill 1935 Genus Cerastes (Horned Vipers)   Cerastes cerastes Sahara Horned Viper Berre 1989 Corkill 1935 [as cornutus] Osman & El Sir 1988 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Werner 1908 [as cornutus]   Cerastes vipera Sahara Sand Viper Abukashawa, Papenfuss & Alkhedir 2018 Sindaco, Venchi & Grieco 2013 Genus Echis (Carpet Vipers)   Echis coloratus Painted Carpet Viper Spawls & Branch 1995   Echis leucogaster White-bellied Carpet Viper Berre 1989   Echis pyramidum North East African Carpet Viper Broadley 1994 Colley 1946 [as E. carinatus] Corkill 1935 [as E. carinatus] Hughes 1976 Osman & El Sir 1988 Pook, Joger, Stumpel & Wuster 2009 Spawls & Branch 1995 Werner 1908 [as E. carinatus]

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa gehafie Eritrean Helmeted Turtle Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelomedusa schweinfurthi Central African Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelomedusa subrufa Common Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Bour 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Werner 1924  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Kinixys (Hinge-back Tortoises)   Kinixys belliana Central African Hinge-back Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 McCord & Joseph-Ouni 2003 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cyclanorbis senegalensis Senegal Flapshell Turtle McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021