Reptiles of South Sudan    Bibliography of the reptiles of South Sudan Taxonomic history of the reptiles of South Sudan Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Fuchs, Mertens & Wermuth 1974 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   EX Mecistops leptorhynchus Central African Slender-snouted Crocodile Shirley, Carr, Nestler, Vliet & Brochu 2018 [as cataphractus]

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama hartmanni Dongola Agama Wagner & Bauer 2011   Agama picticauda Ada Foah Agama Boehme, Wagner, Malonza, Loetters & Koehler 2005 Haxaire & Ineich 1999 [as agama] Loveridge 1957 [as agama] Werner 1908 [as colonorum]   Agama rueppelli Arboreal Agama Drewes 1972 Wagner & Bauer 2011  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo gracilis Graceful Chameleon Friis & Rasmussen 1981 Haxaire & Ineich 1999 Werner 1908   Chamaeleo laevigatus Smooth Chameleon Broadley & Howell 1991 Glaw 2015 Werner 1908 Genus Trioceros (Equatorial Chameleons)   Trioceros bitaeniatus Two-striped Chameleon Boehme & Klaver 1980 Broadley & Howell 1991 Friis & Rasmussen 1981 Glaw 2015 Haxaire & Ineich 1999 Klaver & Boehme 1997   Trioceros conirostratus Sudanese Unicorn Chameleon Glaw 2015 Hughes, Blackburn, Wilber & Behangana 2018 Tilbury 1998 Tilbury 2010   Trioceros ellioti Montane Side-striped Chameleon Boehme & Klaver 1980 Broadley & Howell 1991 Friis & Rasmussen 1981 Glaw 2015 Haxaire & Ineich 1999 Klaver & Boehme 1997 Tilbury 2010   * Trioceros kinetensis Mount Kineti Chameleon Boehme & Klaver 1980 Glaw 2015 Klaver & Boehme 1997 Schmidt 1943 Tilbury 2010   Trioceros rudis Ruwenzori Side-striped Chameleon Glaw 2015 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   Ancylodactylus dickersonae Ituri Four-lined Forest Gecko Friis & Rasmussen 1981 [as africana] Perret 1986 Spawls, Beraduccii & Malonza 2021 Spawls, Howell, Hinkel & Menegon 2018 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Drewes 1972 Haxaire & Ineich 1999 Loveridge 1947 Werner 1908   Hemidactylus lanzai Lake Chew Bahir Half-toed Gecko Drewes 1972 [as H. isolepis] Smid, Mazuch, Novakova, Modry, Malonza, Elmi, Carranza & Moravec 2020 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Loveridge 1947 Werner 1908   Lygodactylus sudanensis Sudan Dwarf Gecko Loveridge 1947 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus sthenodactylus Elegant Thin-toed Gecko Spawls, Howell, Hinkel & Menegon 2018  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987  Family Lacertidae (Typical Lizards) Genus Adolfus (African Forest Lizards)   Adolfus africanus Multi-scaled Forest Lizard Friis & Rasmussen 1981 Haxaire & Ineich 1999 Koehler, Wagner, Visser & Boehme 2003 Spawls, Howell, Hinkel & Menegon 2018 Genus Heliobolus (Bushveld Lizards)   Heliobolus spekii East African Bushveld Lizard Drewes 1972 Lanza 1990 Loveridge 1957 Spawls, Howell, Hinkel & Menegon 2018 Werner 1908 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Drewes 1972  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Joger 1984 Lanza 1990  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides bottegi Ethiopian Cylindrical Skink Drewes 1972 Greenbaum 2005 Greenbaum, Campbell & Raxworthy 2006 Genus Leptosiaphos (Slender Leaf-litter Skinks)   Leptosiaphos aloysiisabaudiae Uganda Five-toed Skink Broadley 1989 Schmidt 1943 [as weberi] Schmitz, Ineich & Chirio 2005 Spawls, Howell, Hinkel & Menegon 2018   Leptosiaphos kilimensis Kilimanjaro Five-toed Skink Ullenbruch & Boehme 2017 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis brevicollis Short-necked Skink Drewes 1972 Haxaire & Ineich 1999   Trachylepis maculilabris Common Speckled-lipped Skink Haxaire & Ineich 1999 Loveridge 1957   Trachylepis megalura Grass-top Skink Haxaire & Ineich 1999   Trachylepis perrotetii Orange-flanked Skink Werner 1908 [as Mabuia mongallensis]   Trachylepis quinquetaeniata African Five-lined Skink Broadley & Bauer 1998 Drewes 1972 Friis & Rasmussen 1981 Haxaire & Ineich 1999 Koppetsch 2020 Werner 1908   Trachylepis striata Eastern Striped Skink Broadley & Howell 1991 Broadley 1977 Broadley 2000 Drewes 1972 Lanza 1990 Werner 1908   Trachylepis varia Savanna Variable Skink Schmidt 1943   Trachylepis wingatii Sudan Grass Skink Koppetsch 2020 Werner 1908  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bayless 2002   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Werner 1908 [as V. ocellatus]   Varanus niloticus Nile Monitor Bayless 2002 Werner 1908

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx colubrinus East African Sand Boa Owen 1956 Werner 1908 [as E. thebaicus]  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis degeni Yellow-flanked Herald Snake Largen & Rasmussen 1993 Loveridge 1957 Pitman 1974 Werner 1908 [as Leptodira attarensis]   Crotaphopeltis hotamboeia Common Herald Snake Corkill 1935 Haxaire & Ineich 1999 Loveridge 1955 Werner 1908 Genus Dasypeltis (African Egg Eaters)   Dasypeltis atra Forest Egg Eater Bates & Broadley 2018 Broadley & Howell 1991 Friis & Rasmussen 1981 Haxaire & Ineich 1999 Largen & Rasmussen 1993 Pitman 1974   Dasypeltis confusa Diamondback Egg Eater Bates & Broadley 2018 Chippaux & Jackson 2019 Trape, Mediannikov & Trape 2012   Dasypeltis scabra Common African Egg Eater Bates & Broadley 2018 Broadley 1983 Chippaux & Jackson 2019 Loveridge 1955 Loveridge 1957 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa weileri Grey Nocturnal Tree Snake Rasmussen 1993 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) Haxaire & Ineich 1999 [as D. t. typus] Loveridge 1955 [as D. typus] Spawls & Branch 1995 [as D. typus] Werner 1908 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Loveridge 1955   Grayia tholloni Nandi Water Snake Corkill 1935 Largen & Rasmussen 1993 Owen 1956 Pitman 1974 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Ullenbruch & Boehme 2017 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Friis & Rasmussen 1981 Haxaire & Ineich 1999 Loveridge 1955 Pitman 1974   Meizodon regularis East African Crowned Smooth Snake Schatti 1985   Meizodon semiornatus Semi-ornate Smooth Snake Corkill 1935 Lanza 1990 Largen & Rasmussen 1993 Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Corkill 1935 Loveridge 1955 Loveridge 1958 Werner 1908 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Corkill 1935 [as Chlorophis emini] Werner 1908 [as Chlorophis emini]   Philothamnus battersbyi Northeastern Green Snake Loveridge 1955 Loveridge 1958 Pitman 1974   Philothamnus bequaerti Uganda Green Snake Hughes 1985 Largen & Rasmussen 1993 Loveridge 1955 [as i. irregularis] Loveridge 1958 [as i. irregularis] Werner 1908 [as Chlorophis irregularis]   Philothamnus heterolepidotus Slender Green Snake Broadley & Howell 1991 Loveridge 1955 Loveridge 1957 Loveridge 1958 Pitman 1974   Philothamnus semivariegatus Variegated Bush Snake Corkill 1935 Loveridge 1955 Loveridge 1958 Pitman 1974 Werner 1908 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps florulentus Egyptian Whip Snake Loveridge 1955 Pitman 1974 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Loveridge 1955 Pitman 1974   Scaphiophis raffreyi Ethiopian Hook-nosed Snake Broadley 1994 Haxaire & Ineich 1999 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus obtusus Egyptian Cat Snake Chippaux & Jackson 2019 Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Haxaire & Ineich 1999 [as dhara] Loveridge 1955 Spawls, Howell, Hinkel & Menegon 2018 [as dhara] Spawls, Malonza & Beraduccii 2021 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Broadley & Wallach 2002 Broadley 2001 Owen 1956 Genus Thrasops (Black Tree Snakes)   Thrasops jacksonii Common Black Tree Snake Broadley & Wallach 2002 Friis & Rasmussen 1981 Haxaire & Ineich 1999 Ullenbruch & Boehme 2017 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas adamanteus Eastern Powdered Cat Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Ullenbruch & Boehme 2017 [as pulverulenta]   Toxicodryas vexator Eastern Black-and-Yellow Tree Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Loveridge 1955 [as blandingii] Pitman 1974 [as blandingii]  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Chippaux & Jackson 2019 Owen 1956 Pitman 1974 Spawls & Branch 1995   Dendroaspis polylepis Black Mamba Haakansson & Madsen 1983 Loveridge 1955 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Hinkel & Menegon 2018 Genus Elapsoidea (African Garter Snakes)   Elapsoidea laticincta Sudanese Garter Snake Broadley 1971 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Jakobsen 1997 Loveridge 1955 Pitman 1974 Spawls & Branch 1995   Elapsoidea loveridgei Northeastern Garter Snake Jakobsen 1997 Genus Naja (Typical Cobras)   Naja ashei East African Spitting Cobra Spawls, Howell, Hinkel & Menegon 2018   Naja haje Egyptian Cobra Corkill 1935 Haxaire & Ineich 1999 Loveridge 1955 Spawls & Branch 1995 Trape, Chirio, Broadley & Wuster 2009   Naja nigricollis Black-necked Spitting Cobra Corkill 1935 Loveridge 1955 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Werner 1908   Naja pallida Eastern Pallid Spitting Cobra Largen & Rasmussen 1993 Spawls & Branch 1995 Spawls, Howell, Hinkel & Menegon 2018   Naja subfulva Brown Forest Cobra Flower 1931 [as N. melanoleuca] Loveridge 1955 [as N. melanoleuca] Pitman 1974 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Ullenbruch & Boehme 2017 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus jacksonii Sudan Centipede Eater Lanza 1990 Largen & Rasmussen 1993 Loveridge 1955 Pitman 1974 Wallach, Williams & Boundy 2014   Aparallactus lunulatus Reticulated Centipede Eater Loveridge 1940 Loveridge 1955 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Wallach, Williams & Boundy 2014 Werner 1908 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake Rasmussen 2005 Spawls & Branch 1995   Atractaspis fallax False Stiletto Snake Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake Friis & Rasmussen 1981 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haxaire & Ineich 1999 Loveridge 1955 Schmidt 1943 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis phillipsi Sudanese Stiletto Snake Corkill 1935 [as A. microlepidota] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as A. m. microlepidota] Loveridge 1955 [as A. m. microlepidota] Loveridge 1957 [as A. m. microlepidota] Pitman 1974 [as A. m. microlepidota] Spawls & Branch 1995 [as A. microlepidota]   Atractaspis watsoni Sokoto Stiletto Snake Trape, Mane & Ineich 2006 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Broadley 1983 Corkill 1935 Haxaire & Ineich 1999 Pitman 1974   Boaedon lineatus Lined House Snake Loveridge 1955 Werner 1908   Boaedon perisilvestris Central African Savanna House Snake Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020 Genus Chilorhinophis (Black-and-Yellow Burrowing Snakes)   Chilorhinophis butleri Sudan Black-and-Yellow Burrowing Snake Loveridge 1958 Spawls, Howell, Drewes & Ashe 2002 Werner 1908 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Hughes 2004 Loveridge 1955 Pitman 1974 Werner 1908 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis hildebrandtii Kenyan Bark Snake Broadley & Hughes 2000   Hemirhagerrhis kelleri Stripe-bellied Bark Snake Broadley & Hughes 2000   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Hughes 2000 Broadley 1983 Owen 1956 Werner 1908 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Owen 1956 Genus Kladirostratus (Striped Beaked Snakes)   Kladirostratus togoensis Western Striped Beaked Snake Chirio & Ineich 1991 Keates, Conradie, Greenbaum & Edwards 2019 Genus Limaformosa (Venus File Snakes)   Limaformosa savorgnani Central African Forest File Snake Lanza & Broadley 2014 Lanza 1990 Loveridge 1957 Owen 1956 Pitman 1974 Werner 1908 [as Simocephalus butleri] Genus Lycophidion (African Wolf Snakes)   Lycophidion depressirostre Flat-snouted Wolf Snake Broadley & Hughes 1993 Broadley 1996 Chippaux & Jackson 2019   Lycophidion irroratum Western Forest Wolf Snake Broadley & Hughes 1993   Lycophidion jacksoni Kilimanjaro Wolf Snake Broadley & Hughes 1993 Broadley 1996 Laurent 1968 Loveridge 1955 [as L. c. capense] Trape 2021   Lycophidion ornatum Ornate Wolf Snake Broadley & Hughes 1993 Broadley 1996 Laurent 1968 Genus Polemon (African Snake Eaters)   Polemon christyi Eastern Snake Eater Owen 1956 Genus Prosymna (Shovel-snout Snakes)   Prosymna ambigua Central African Shovel-snout Broadley & Howell 1991 Broadley 1980 Chippaux & Jackson 2019 Largen & Rasmussen 1993 Loveridge 1955 Loveridge 1957 Loveridge 1958 Pitman 1974   Prosymna greigerti Sudan Savanna Shovel-snout Broadley 1980 Loveridge 1955 [as P. meleagris] Loveridge 1958 [as P. laurenti] Papenfuss 1969 [as P. laurenti] Pitman 1974 [as P. laurenti] Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis mossambicus East African Olive Grass Snake Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis rukwae Lake Rukwa Sand Snake Largen & Rasmussen 1993 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis sudanensis Sudan Sand Snake Corkill 1935 [as P. subtaeniatus] Loveridge 1940 Loveridge 1955 Loveridge 1957 Pitman 1974   Psammophis tanganicus Tanganyika Sand Snake Largen & Rasmussen 1993 Genus Psammophylax (Skaapstekers)   Psammophylax multisquamis Kenyan Striped Skaapsteker Owen 1956 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis rostratus Rufous Beaked Snake Broadley & Howell 1991 Broadley 1983 Chirio & Ineich 1991 Lanza 1990 Largen & Rasmussen 1993 Loveridge 1955 Pitman 1974   Rhamphiophis rubropunctatus Red-spotted Beaked Snake Chirio & Ineich 1991 Lanza 1990 Largen & Rasmussen 1993 Loveridge 1955 Pitman 1974 Werner 1908  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops emini Karagwe Thread Snake Broadley & Wallach 2007 Loveridge 1955 Loveridge 1957 Pitman 1974 Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis braccianii Eritrean Mountain Thread Snake Broadley & Wallach 2007 Loveridge 1955 [as cairi]  Family Micrelapidae (Short-jawed Rear-fanged Snakes) Genus Micrelaps (Afro-Asian Black-headed Snakes)   Micrelaps vaillanti Somali Black-headed Snake Lanza 1990 [as M. boettgeri] Loveridge 1955 [as M. boettgeri] Pitman 1974 [as M. boettgeri] Rasmussen 2002  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Owen 1956 Pitman 1974 Werner 1908   Python sebae Northern Rock Python Broadley 1984 Owen 1956 Spawls & Branch 1995 Werner 1908  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops brevis Somali Giant Blind Snake Broadley & Wallach 2009 Hahn 1980 Largen & Rasmussen 1993 Loveridge 1955 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Werner 1908 [as schlegelii]   Afrotyphlops lineolatus Lineolate Blind Snake Broadley & Wallach 2009 Hahn 1980 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Broadley & Wallach 2009 Corkill 1935 Loveridge 1955 Schmidt 1943 Wallach, Williams & Boundy 2014 Werner 1908 Genus Letheobia (African Gracile Blind Snakes)   * Letheobia toritensis Torit Gracile Blind Snake Broadley & Wallach 2007 Loveridge 1955 [as pallida]  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   Atheris squamigera Southern Green Bush Viper Ullenbruch & Boehme 2017 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Corkill 1935 Haxaire & Ineich 1999 Owen 1956 Spawls & Branch 1995 Werner 1908   Bitis gabonica Gaboon Adder Broadley & Howell 1991 Broadley 1983 Loveridge 1957 Owen 1956 Pitman 1974 Spawls & Branch 1995   Bitis nasicornis African Nose-horned Viper Broadley & Howell 1991 Friis & Rasmussen 1981 Haxaire & Ineich 1999 Loveridge 1955 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus lichtensteinii Forest Night Adder Friis & Rasmussen 1981 Haxaire & Ineich 1999 Loveridge 1955 Pitman 1974   Causus maculatus West African Night Adder Haxaire & Ineich 1999 Largen & Rasmussen 1993 Spawls & Branch 1995   Causus resimus Green Night Adder Corkill 1935 Loveridge 1955 Pitman 1974 Spawls & Branch 1995 Werner 1908   Causus rhombeatus Rhombic Night Adder Corkill 1935 Loveridge 1955 Spawls & Branch 1995 Genus Echis (Carpet Vipers)   Echis pyramidum North East African Carpet Viper Drewes 1972 Pitman 1974 [as E. carinatus leakeyi] Spawls & Branch 1995

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa schweinfurthi Central African Helmeted Turtle Drewes 1972 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios chapini Central African Mud Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys belliana Central African Hinge-back Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Kinixys erosa Forest Hinge-back Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Kinixys nogueyi Western Savanna Hinge-back Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wermuth & Mertens 1977  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 Demaya, Benansio, Lado, Diagne, Dendi & Luiselli 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cyclanorbis senegalensis Senegal Flapshell Turtle McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021