Reptiles of the Central African Republic    Bibliography of the reptiles of the Central African Republic Taxonomic history of the reptiles of the Central African Republic Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca leucura White-tailed Worm Lizard Chirio & Ineich 2006 Gans 1987 Gans 2005 Joger 1990 Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis kabindae Cabinda Spade-snouted Worm Lizard Santos 2013

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Chirio & Ineich 2006 [as niloticus] Fuchs, Mertens & Wermuth 1974 Joger 1990 [as niloticus] Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   Mecistops leptorhynchus Central African Slender-snouted Crocodile Chirio & Ineich 2006 [as cataphractus] Joger 1990 [as cataphractus] Shirley, Carr, Nestler, Vliet & Brochu 2018 Trape, Trape & Chirio 2012 [as cataphractus] Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Chirio & Ineich 2006 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama doriae Nigerian Agama Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Moody & Boehme 1984 Trape, Trape & Chirio 2012   Agama gracilimembris Lesser Ground Agama Chirio & Ineich 2006 Grandison 1968 Joger 1990 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Wagner & Bauer 2011   Agama picticauda Ada Foah Agama Chirio & Ineich 2006 [as agama] Jacobsen 2009 [as agama] Joger 1990 [as agama] Trape, Trape & Chirio 2012 [as agama]  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo africanus Sahel Chameleon Chirio & Ineich 2006 Glaw 2015 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo gracilis Graceful Chameleon Chirio & Ineich 2006 Glaw 2015 Jacobsen 2009 Joger 1990 Klaver & Boehme 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo laevigatus Smooth Chameleon Glaw 2015 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Chirio & Ineich 2006 Tilbury 2010 Genus Rhampholeon (African Leaf Chameleons)   Rhampholeon spectrum Spectral Leaf Chameleon Chirio & Ineich 2006 Glaw 2015 Joger 1990 Tilbury 2010 Trape, Trape & Chirio 2012 Genus Trioceros (Equatorial Chameleons)   Trioceros cristatus Crested Chameleon Chirio & Ineich 2006 Glaw 2015 Joger 1990 Klaver & Boehme 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Trioceros oweni Central African Three-horned Chameleon Chirio & Ineich 2006 Joger 1990 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Chirio & Ineich 2006 Chirio & LeBreton 2007 Jacobsen 2009 Joger 1990 [as brookii] Trape, Trape & Chirio 2012   Hemidactylus longicephalus Long-headed Half-toed Gecko Chirio & Ineich 2006 Joger 1990   Hemidactylus mabouia Cosmopolitan House Gecko Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Hemidactylus pseudomuriceus Azagny Half-toed Gecko Koppetsch & Boehme 2017   Hemidactylus richardsonii Central African Half-toed Gecko Chirio & Ineich 2006 Trape, Trape & Chirio 2012 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus depressus Spotted Dwarf Gecko Chirio & Ineich 2006   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987 Chirio & Ineich 2006 Joger 1990 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus guineensis Guinea Fringe-toed Lizard Chirio & Ineich 2006 Trape, Trape & Chirio 2012 Genus Adolfus (African Forest Lizards)   Adolfus africanus Multi-scaled Forest Lizard Chirio & Ineich 2006 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Chirio & Ineich 2006 Joger 1990 Szczerbak 1975 Trape, Trape & Chirio 2012 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Chirio & Ineich 2006 Trape, Trape & Chirio 2012 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Chirio & Ineich 2006 Trape, Trape & Chirio 2012 Genus Poromera (African Swamp Lizard)   Poromera fordii African Swamp Lizard Chirio & Ineich 2006  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Chirio & Ineich 2006 Joger 1984 Joger 1990 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola ephippiata African Wall Gecko Chirio & Ineich 2006 Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Feylinia (African Snake Skinks)   Feylinia currori Western Forest Snake Skink Brygoo & Roux-Esteve 1983 Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Feylinia elegans Elegant Snake Skink Chirio & Ineich 1992 Chirio & Ineich 2006   Feylinia grandisquamis Cameroon Snake Skink Brygoo & Roux-Esteve 1983 Chirio & Ineich 2006 Joger 1990   Feylinia macrolepis Congolese Snake Skink Chirio & Ineich 2006 Genus Lacertaspis (Central African Leaf-litter Skinks)   Lacertaspis reichenowii Cameroon Mountain Leaf-Litter Skink Chirio & Ineich 2006 Schmitz, Ineich & Chirio 2005 Genus Melanoseps (Black Limbless Skinks)   Melanoseps occidentalis Western Limbless Skink Chirio & Ineich 2006 Trape, Trape & Chirio 2012 Genus Mochlus (African Supple Skinks)   Mochlus fernandi Western Red-flanked Skink Trape, Trape & Chirio 2012   Mochlus striatus Central African Red-flanked Skink Wagner, Boehme, Pauwels & Schmitz 2009   Mochlus sundevallii Common Savanna Supple Skink Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Genus Panaspis (African Snake-eyed Skinks)   Panaspis breviceps Short-headed Snake-eyed Skink Chirio & Ineich 2006 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012   Panaspis togoensis Togo Snake-eyed Skink Chirio & Ineich 2006 Joger 1990 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Trachylepis buettneri Togo Savanna Skink Chirio & Ineich 2006 Joger 1990 Trape, Trape & Chirio 2012   Trachylepis maculilabris Common Speckled-lipped Skink Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Trachylepis makolowodei Belemboke Skink Chirio & Ineich 2006 [as "Trachylepis sp. 2"] Chirio, Ineich, Schmitz & LeBreton 2008 Gvozdik, Dolinay, Zassi-Boulou & Babangenge 2018   * Trachylepis pendeana Pende River Skink Chirio & Ineich 2006 Ineich & Chirio 2000   Trachylepis perrotetii Orange-flanked Skink Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Trachylepis polytropis Greater Forest Skink Chirio & Ineich 2006 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Koppetsch 2020 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Chirio & Ineich 2006 Trape, Trape & Chirio 2012   Varanus stellatus African Forest Monitor Boehme & Ziegler 1997 [as V. ornatus] Boehme & Ziegler in Pianka, King & King (eds.) 2004 [as V. ornatus] Chirio & Ineich 2006 [as V. ornatus] Jacobsen 2009 [as V. ornatus] Trape, Trape & Chirio 2012 [as V. ornatus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Chirio & Ineich 2006 Joger 1990 Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Chirio & Ineich 2006  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis degeni Yellow-flanked Herald Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Rasmussen, Chirio & Ineich 2000   Crotaphopeltis hippocrepis West African Herald Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Rasmussen, Chirio & Ineich 2000   Crotaphopeltis hotamboeia Common Herald Snake Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Rasmussen, Chirio & Ineich 2000 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Bates & Broadley 2018 Chirio & LeBreton 2007 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Bates & Broadley 2018 Chirio & Ineich 2006 Joger 1990 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa duchesnei Blue-tailed Nocturnal Tree Snake Chirio & Ineich 2006 Joger 1990 Rasmussen 1989   Dipsadoboa underwoodi Cameroon Nocturnal Tree Snake Broadley & Cassidy 2011   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Chirio & Ineich 2006 Joger 1990   Dipsadoboa viridis Green Nocturnal Tree Snake Chirio & Ineich 2006   Dipsadoboa weileri Grey Nocturnal Tree Snake Chirio & Ineich 2006 Rasmussen 1993 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) Chirio & Ineich 2006 [as D. typus] David & Ineich 1999 [as D. typus] Joger 1990 [as D. typus] Spawls & Branch 1995 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia caesar Long-tailed Water Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Grayia ornata Ornate Water Snake Chirio & Ineich 2006   Grayia smythii Equatorial Water Snake Chirio & Ineich 2006 Jacobsen 2009 Joger 1990   Grayia tholloni Nandi Water Snake Chirio & Ineich 2006 Joger 1990 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Chirio & Ineich 2006 Joger 1990   Hapsidophrys smaragdinus Common Emerald Snake Chirio & Ineich 2006 Joger 1990 Genus Hydraethiops (Central African Water Snakes)   Hydraethiops melanogaster Black-bellied Water Snake Chirio & Ineich 2006 Joger 1990 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Chirio & Ineich 2006 Joger 1990   Meizodon regularis East African Crowned Smooth Snake Chirio & Ineich 2006 Jacobsen 2009 Genus Natriciteres (African Marsh Snakes)   Natriciteres fuliginoides Collared Marsh Snake Chirio & Ineich 2006 Joger 1990   Natriciteres olivacea Olive Marsh Snake Chirio & Ineich 1992 Chirio & Ineich 2006 Joger 1990   Natriciteres variegata Variable Marsh Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Jacobsen 2009 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Chirio & Ineich 2006 Joger 1990   Philothamnus bequaerti Uganda Green Snake Chirio & Ineich 2006 Jacobsen 2009 Joger 1990   Philothamnus carinatus Barred Green Snake Chirio & Ineich 2006 Joger 1990   Philothamnus heterodermus Emerald Green Snake Chirio & Ineich 2006 Joger 1990   Philothamnus heterolepidotus Slender Green Snake Chirio & Ineich 2006 Joger 1990   Philothamnus hughesi Congolese Green Snake Chirio & Ineich 2006 Trape & Roux-Esteve 1990   Philothamnus irregularis Irregular Green Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Philothamnus nitidus Cameroon Green Snake Broadley & Howell 1991 Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Philothamnus ornatus Ornate Green Snake Chirio & LeBreton 2007   Philothamnus semivariegatus Variegated Bush Snake Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Chirio & Ineich 2006 Joger 1990   Rhamnophis batesii Spotted Dagger-tooth Tree Snake Chirio & Ineich 2006 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Chirio & Ineich 2006 Joger 1990 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus tripolitanus North African Cat Snake Chirio & Ineich 2006 [as obtusus] Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Sindaco, Venchi & Grieco 2013 Wilms, Wagner & Spawls 2021   Telescopus variegatus Variable Cat Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Chirio & Ineich 2006 Joger 1990 Spawls & Branch 1995 Genus Thrasops (Black Tree Snakes)   Thrasops flavigularis Yellow-throated Black Tree Snake Chirio & Ineich 2006   Thrasops jacksonii Common Black Tree Snake Broadley & Wallach 2002 Chirio & Ineich 2006 Joger 1990 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas adamanteus Eastern Powdered Cat Snake Chirio & Ineich 2006 [as pulverulenta] Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Joger 1990 [as pulverulenta]   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Chirio & Ineich 2006 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Jacobsen 2009 Joger 1990 Spawls & Branch 1995   Toxicodryas vexator Eastern Black-and-Yellow Tree Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Chippaux & Jackson 2019 Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 Spawls & Branch 1995   Dendroaspis polylepis Black Mamba Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 Spawls & Branch 1995 Genus Elapsoidea (African Garter Snakes)   Elapsoidea laticincta Sudanese Garter Snake Broadley 1971 Chippaux & Jackson 2019 Chirio & Ineich 2006 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Jakobsen 1997 Joger 1990 Spawls & Branch 1995   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1971 Chirio & Ineich 2006 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Joger 1990 Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja annulata Ringed Water Cobra Chippaux & Jackson 2019 Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 Spawls & Branch 1995   Naja haje Egyptian Cobra Sindaco, Venchi & Grieco 2013 Trape, Chirio, Broadley & Wuster 2009   Naja melanoleuca Central African Forest Cobra Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 Spawls & Branch 1995 Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja multifasciata Burrowing Cobra Chirio & Ineich 2006 David & Ineich 1999   Naja nigricollis Black-necked Spitting Cobra Chirio & Ineich 2006 Joger 1990 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Naja subfulva Brown Forest Cobra Chirio & Ineich 2006 Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018 Genus Pseudohaje (Tree Cobras)   Pseudohaje goldii Central African Tree Cobra Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 Spawls & Branch 1995  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Chirio & Ineich 2006 Ineich & Chirio 1992 Joger 1990 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus lunulatus Reticulated Centipede Eater Chirio & Ineich 2006 Joger 1990 Wallach, Williams & Boundy 2014   Aparallactus modestus Western Forest Centipede Eater Chirio & Ineich 2006 Joger 1990 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis boulengeri Central African Stiletto Snake Chirio & Ineich 2006 David & Ineich 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis corpulenta Fat Stiletto Snake Chirio & Ineich 2006 David & Ineich 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis reticulata Reticulate Stiletto Snake Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis watsoni Sokoto Stiletto Snake Chirio & Ineich 2006 Trape, Mane & Ineich 2006 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Chirio & Ineich 2006 Jacobsen 2009 Joger 1990   Boaedon lineatus Lined House Snake Chirio & Ineich 2006 Joger 1990   Boaedon olivaceus Olive House Snake Chirio & Ineich 2006 Joger 1990   Boaedon paralineatus Central African Lined House Snake Trape & Mediannikov 2016   Boaedon perisilvestris Central African Savanna House Snake Trape & Mediannikov 2016 Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020   Boaedon subflavus Yellow House Snake Trape & Mediannikov 2016 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus lineatus Red-and-Black Striped Snake Chirio & Ineich 2006 Joger 1990 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Chirio & Ineich 2006 Joger 1990 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Chirio & Ineich 2006 Hughes 2004 Joger 1990   Dromophis praeornatus Ornate Olympic Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Hughes & Barry 1969 Jacobsen 2009 Joger 1990 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis brussauxi Eastern Forest Dwarf File Snake Chirio & LeBreton 2007 Ineich, Fretey, Colyn & Barriere 2007   Gonionotophis grantii Savanna Dwarf File Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Hughes 2000 Chirio & Ineich 2006 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Chirio & Ineich 2006 Joger 1990 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Chirio & Ineich 2006 Joger 1990   Limaformosa savorgnani Central African Forest File Snake Chirio & Ineich 2006 Joger 1990 [as capensis] Lanza & Broadley 2014 Lanza 1990 Genus Lycophidion (African Wolf Snakes)   Lycophidion chirioi Sangba Wolf Snake Chirio & Ineich 2006 [as L. capense] Joger 1990 [as L. capense] Trape 2021   Lycophidion depressirostre Flat-snouted Wolf Snake Chippaux & Jackson 2019 Chirio & Ineich 2006   Lycophidion irroratum Western Forest Wolf Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Lycophidion jacksoni Trape 2021   Lycophidion laterale Masked Wolf Snake Chirio & Ineich 2006   Lycophidion multimaculatum Many-spotted Wolf Snake Chippaux & Jackson 2019   Lycophidion semicinctum Semi-annulated Wolf Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Lycophidion tchadensis Chadian Wolf Snake Chirio & Ineich 2006 [as taylori] Trape 2021 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Chirio & Ineich 2006 Joger 1990   Mehelya stenophthalmus Small-eyed File Snake Chirio & Ineich 2006 Joger 1990 Genus Polemon (African Snake Eaters)   Polemon collaris Fawn-headed Snake Eater Chirio & Ineich 2006 Joger 1990   Polemon gabonensis Gabon Snake Eater Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Polemon notatus African Spotted Snake Eater Chirio & Ineich 2006   Polemon robustus Robust Snake Eater Chippaux & Jackson 2019 Chirio & Ineich 2006 Genus Prosymna (Shovel-snout Snakes)   Prosymna ambigua Central African Shovel-snout Broadley 1980 Chirio & Ineich 2006 Jacobsen 2009 Joger 1990   Prosymna collaris Collared Shovel-snout Chirio & Ineich 2006 Chirio, Ineich, Schmitz & Trape 2011   Prosymna greigerti Sudan Savanna Shovel-snout Chirio & Ineich 2006 Chirio, Ineich, Schmitz & Trape 2011 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis elegans Elegant Sand Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990   Psammophis mossambicus East African Olive Grass Snake Chirio & Ineich 2006 [as phillipsii] Chirio & LeBreton 2007 [as occidentalis] Hughes & Barry 1969 [as phillipsii] Jacobsen 2009 [as phillipsii] Joger 1990 [as phillipsii] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis rukwae Lake Rukwa Sand Snake Chirio & Ineich 2006 [as P. sibilans] Joger 1990 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis sudanensis Sudan Sand Snake Chirio & Ineich 2006 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chirio & Ineich 2006 Joger 1990  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis narirostris Sahel Thread Snake Chirio & Ineich 2006 Jacobsen 2009   Myriopholis occipitalis Kouki Thread Snake Trape & Chirio 2019 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma sundewalli Central African Thread Snake Chippaux & Jackson 2019 Chirio & Ineich 2006 Hahn 1980 Joger 1990 McDiarmid, Campbell & Toure 1999  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Chirio & Ineich 2006 Jacobsen 2009 Joger 1990   Python sebae Northern Rock Python Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops angolensis Golden Blind Snake Chirio & Ineich 2006 Hahn 1980 Joger 1990 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   * Afrotyphlops chirioi Berberati Blind Snake Trape 2019   Afrotyphlops congestus Marbled Blind Snake Chirio & Ineich 2006 Hahn 1980 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops decorosus Cameroon Gracile Blind Snake Broadley & Wallach 2007 Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990 McDiarmid, Campbell & Toure 1999   Afrotyphlops lineolatus Lineolate Blind Snake Chirio & Ineich 2006 Joger 1990 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Chirio & Ineich 2006 Hahn 1980 Jacobsen 2009 Joger 1990 Wallach, Williams & Boundy 2014   Afrotyphlops steinhausi Common Cameroon Blind Snake Broadley & Wallach 2009 Chirio & Ineich 2006 Hahn 1980 Joger 1990 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014 Genus Letheobia (African Gracile Blind Snakes)   * Letheobia debilis Feeble Gracile Blind Snake Broadley & Wallach 2007 Chippaux & Jackson 2019 Chirio & Ineich 2006 Joger 1990 McDiarmid, Campbell & Toure 1999   Letheobia rufescens Oubangui Gracile Blind Snake Broadley & Wallach 2007 Chippaux & Jackson 2019 Chirio & Ineich 2006 Jacobsen 2009 McDiarmid, Campbell & Toure 1999   Letheobia wittei Ubangi Gracile Blind Snake Chirio & Ineich 2006  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   Atheris broadleyi Lipondji Bush Viper Chippaux & Jackson 2019 Chirio & Ineich 2006 Wallach, Williams & Boundy 2014   Atheris squamigera Southern Green Bush Viper Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995   Bitis gabonica Gaboon Adder Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Spawls & Branch 1995   Bitis nasicornis African Nose-horned Viper Chirio & Ineich 2006 David & Ineich 1999 Joger 1990 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus lichtensteinii Forest Night Adder Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 Spawls & Branch 1995   Causus maculatus West African Night Adder Chirio & Ineich 2006 David & Ineich 1999 Jacobsen 2009 Joger 1990 Spawls & Branch 1995   Causus resimus Green Night Adder Chirio & Ineich 2006 David & Ineich 1999 Spawls & Branch 1995   Causus rhombeatus Rhombic Night Adder David & Ineich 1999 Jacobsen 2009 Genus Echis (Carpet Vipers)   Echis pyramidum North East African Carpet Viper Chippaux & Jackson 2019 Chirio & Ineich 2006 David & Ineich 1999   Echis romani Chadian Carpet Viper Chippaux & Jackson 2019 [as ocellatus] Chirio & Ineich 2006 [as ocellatus] David & Ineich 1999 [as ocellatus] Joger 1990 [as ocellatus] Spawls & Branch 1995 [as ocellatus] Trape 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa schweinfurthi Central African Helmeted Turtle Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelomedusa subrufa Common Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Bonin, Devaux & Dupre 2006 Bour 2008 Chirio & Ineich 2006 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Chirio & Ineich 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios chapini Central African Mud Turtle Chirio & Ineich 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios gabonensis Forest Mud Turtle Chirio & Ineich 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios rhodesianus Zambian Mud Turtle Jacobsen 2009   Pelusios subniger East African Black Mud Turtle Jacobsen 2009  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Chirio & Ineich 2006 Joger 1990 Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Kinixys (Hinge-back Tortoises)   Kinixys belliana Central African Hinge-back Tortoise Chirio & Ineich 2006 Jacobsen 2009 Joger 1990 Kindler, Branch, Hofmeyr, Maran, Siroky, Vences, Harvey, Hauswaldt, Schleicher, Stuckas & Fritz 2012 McCord, Joseph-Ouni & Tabaka 2005   Kinixys erosa Forest Hinge-back Tortoise Chirio & Ineich 2006 Joger 1990 Luiselli & Diagne 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys homeana West African Hinge-back Tortoise Chirio & Ineich 2006 Luiselli & Diagne 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Trape, Trape & Chirio 2012   Kinixys nogueyi Western Savanna Hinge-back Tortoise Chirio & LeBreton 2007 Fritz & Havas 2006 Iverson 1992 McCord, Joseph-Ouni & Tabaka 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cyclanorbis senegalensis Senegal Flapshell Turtle Chirio & Ineich 2006 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Cycloderma (Central African Flapshell Turtles)   Cycloderma aubryi Forest Flapshell Turtle Chirio & Ineich 2006 Fritz & Havas 2006 Joger 1990 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Chirio & Ineich 2006 Joger 1990 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012