Reptiles of Uganda    Bibliography of the reptiles of Uganda Taxonomic history of the reptiles of Uganda Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Behangana 2004 Fuchs, Mertens & Wermuth 1974 King & Burke 1989 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wermuth & Mertens 1977 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus osborni Congo Dwarf Crocodile Spawls, Howell, Hinkel & Menegon 2018

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus kiwuensis Kivu Ridgeback Agama Wagner, Butler, Ceriaco & Bauer 2021 Wagner, Greenbaum & Bauer 2012 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018   Acanthocercus ugandaensis Uganda Ridgeback Agama Klausewitz 1957 Wagner, Butler, Ceriaco & Bauer 2021 Wagner, Greenbaum & Bauer 2012 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018 Wermuth 1967 Genus Agama (Typical Agamas)   Agama finchi West Kenyan Rainbow Agama Behangana 2004 [as agama] Spawls, Howell, Hinkel & Menegon 2018 Wagner, Freund, Modry, Schmitz & Boehme 2011   Agama lionotus Kenyan Rock Agama Spawls, Howell, Hinkel & Menegon 2018   Agama rueppelli Arboreal Agama Spawls, Howell, Hinkel & Menegon 2018  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Glaw 2015 Klaver & Boehme 1997 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   Chamaeleo gracilis Graceful Chameleon Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   Chamaeleo laevigatus Smooth Chameleon Behangana 2004 Glaw 2015 Klaver & Boehme 1997 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010 Genus Kinyongia (East African Dwarf Chameleons)   Kinyongia carpenteri Ruwenzori Helmeted Chameleon Glaw 2015 Klaver & Boehme 1997 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   * Kinyongia tolleyae Kigezi Highlands Dwarf Chameleon Glaw 2015 [as adolfifriderici] Greenbaum, Tolley, Joma & Kusamba 2012 [as adolfifriderici] Hughes, Kusamba, Behangana & Greenbaum 2017 Klaver & Boehme 1997 [as adolfifriderici] Spawls, Howell, Drewes & Ashe 2002 [as adolfifriderici] Spawls, Howell, Hinkel & Menegon 2018 [as adolfifriderici] Tilbury 2010 [as adolfifriderici] Vonesh 1998 [as adolfifriderici]   Kinyongia xenorhina Ruwenzori Plate-nosed Chameleon Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010 Genus Rhampholeon (African Leaf Chameleons)   Rhampholeon boulengeri Central African Leaf Chameleon Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010 Vonesh 1998 Genus Trioceros (Equatorial Chameleons)   Trioceros bitaeniatus Two-striped Chameleon Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   Trioceros conirostratus Sudanese Unicorn Chameleon Hughes, Blackburn, Wilber & Behangana 2018 Spawls, Howell, Hinkel & Menegon 2018   Trioceros ellioti Montane Side-striped Chameleon Broadley & Howell 1991 Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010 Vonesh 1998   Trioceros hoehnelii High-casqued Chameleon Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   Trioceros ituriensis Ituri Forest Chameleon Glaw 2015 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Trioceros johnstoni Ruwenzori Three-horned Chameleon Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010   Trioceros rudis Ruwenzori Side-striped Chameleon Broadley & Howell 1991 Glaw 2015 Klaver & Boehme 1997 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Chamaesaura (African Grass Lizards)   Chamaesaura tenuior East African Highland Grass Lizard Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   Ancylodactylus dickersonae Ituri Four-lined Forest Gecko Malonza & Bauer 2022 Perret 1986 Spawls, Beraduccii & Malonza 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Ancylodactylus elgonensis Elgon Forest Gecko Malonza & Bauer 2022 Perret 1986 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wagner, Safari & Chenga 2021   Ancylodactylus quattuorseriatus Kivu Four-lined Forest Gecko Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wagner, Safari & Chenga 2021 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Behangana 2004 [as mabouia brooki] Loveridge 1947 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 [as brooki] Spawls, Howell, Hinkel & Menegon 2018   Hemidactylus mabouia Cosmopolitan House Gecko Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Hemidactylus richardsonii Central African Half-toed Gecko Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Loveridge 1947 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Lygodactylus keniensis Kenya Dwarf Gecko Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Spawls, Howell, Hinkel & Menegon 2018   Gerrhosaurus intermedius Eastern Black-lined Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Spawls, Howell, Drewes & Ashe 2002 [as nigrolineatus] Spawls, Howell, Hinkel & Menegon 2018 [as nigrolineatus]  Family Lacertidae (Typical Lizards) Genus Adolfus (African Forest Lizards)   Adolfus africanus Multi-scaled Forest Lizard Arnold 1989 Greenbaum, Villanueva, Kusamba, Aristote & Branch 2011 Koehler, Wagner, Visser & Boehme 2003 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Adolfus jacksoni East African Forest Lizard Arnold 1989 Broadley & Howell 1991 Greenbaum, Beer, Hughes, Wagner, Anderson, Villanueva, Malonza, Kusumba, Muninga, Aristote & Branch 2018 Greenbaum, Villanueva, Kusamba, Aristote & Branch 2011 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Adolfus masavaensis Masava Meadow Lizard Arnold 1989 [as alleni] Greenbaum, Villanueva, Kusamba, Aristote & Branch 2011 [as alleni] Loveridge 1957 [as alleni] Wagner, Greenbaum, Malonza & Branch 2014 Genus Congolacerta (Congolese Lizards)   Congolacerta vauereselli Sparse-scaled Forest Lizard Arnold 1989 Broadley & Howell 1991 Greenbaum, Villanueva, Kusamba, Aristote & Branch 2011 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Spawls, Howell, Hinkel & Menegon 2018   Heliobolus spekii East African Bushveld Lizard Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Arnold 1989 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Spawls, Howell, Hinkel & Menegon 2018  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Jirku, Mihalca, Necas & Modry 2010  Family Scincidae (Skinks) Genus Feylinia (African Snake Skinks)   Feylinia currori Western Forest Snake Skink Brygoo & Roux-Esteve 1983 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Leptosiaphos (Slender Leaf-litter Skinks)   Leptosiaphos aloysiisabaudiae Uganda Five-toed Skink Peracca 1907 Schmitz, Ineich & Chirio 2005 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Leptosiaphos graueri Rwanda Five-toed Skink Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Leptosiaphos hackarsi Virunga Four-toed Skink Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Leptosiaphos kilimensis Kilimanjaro Five-toed Skink Spawls, Howell, Hinkel & Menegon 2018   Leptosiaphos meleagris Ruwenzori Four-toed Skink Ineich 2010 [as Panaspis helleri] Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Mochlus (African Supple Skinks)   Mochlus hinkeli Eastern Red-flanked Skink Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 [as fernandi] Wagner, Boehme, Pauwels & Schmitz 2009   Mochlus sundevallii Common Savanna Supple Skink Broadley 1966 Broadley 1994 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 [as M. afer] Genus Panaspis (African Snake-eyed Skinks)   Panaspis wahlbergii Savanna Snake-eyed Skink Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis albilabris African White-lipped Forest Skink Mausfeld & Schmitz 2000 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis bayonii Equatorial Mountain Grass Skink Kingdon & Spawls 2010   Trachylepis brevicollis Short-necked Skink Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis irregularis Alpine Meadow Skink Broadley 1977 Loveridge 1957 Spawls, Howell, Hinkel & Menegon 2018   * Trachylepis loluiensis Lolui Island Skink Kingdon & Spawls 2010 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis maculilabris Common Speckled-lipped Skink Behangana 2004 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Trachylepis margaritifer African Rainbow Skink Broadley & Howell 1991   Trachylepis megalura Grass-top Skink Broadley 2000 Koppetsch 2020 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Trachylepis perrotetii Orange-flanked Skink Chirio & LeBreton 2007 Hoogmoed 1974 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis planifrons Teita Skink Koppetsch 2020 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis quinquetaeniata African Five-lined Skink Behangana 2004 Broadley & Bauer 1998 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Trachylepis striata Eastern Striped Skink Behangana 2004 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Trachylepis varia Savanna Variable Skink Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bayless 2002 Loveridge 1957 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Spawls, Howell, Hinkel & Menegon 2018   Varanus niloticus Nile Monitor Bayless 2002 Behangana 2004 Loveridge 1957

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx colubrinus East African Sand Boa Spawls, Howell, Hinkel & Menegon 2018  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis degeni Yellow-flanked Herald Snake Loveridge 1957 Pitman 1974 Rasmussen 1985 Rasmussen, Chirio & Ineich 2000 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Crotaphopeltis hotamboeia Common Herald Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Dasypeltis (African Egg Eaters)   Dasypeltis atra Forest Egg Eater Bates & Broadley 2018 Hughes 1983 Hughes 1997 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Dasypeltis confusa Diamondback Egg Eater Bates & Broadley 2018 Spawls, Howell, Hinkel & Menegon 2018 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Bates & Broadley 2018 Gans 1959 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Dasypeltis scabra Common African Egg Eater Bates & Broadley 2018 Gans 1959 Hughes 1997 Pitman 1974 Spawls, Howell, Hinkel & Menegon 2018 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Branch, Conradie & Tolley in Branch, Bayliss, Bittencourt-Silva, Conradie, Engelbrecht, Loader, Menegon, Nanvonamuquitxo & Tolley 2019 Chippaux & Jackson 2019 Hughes 1983 Loveridge 1957 Pitman 1974 Rasmussen 1993 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Dipsadoboa weileri Grey Nocturnal Tree Snake Rasmussen 1993 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Dispholidus (Boomslangs)   Dispholidus kivuensis Kivu Boomslang Pitman 1974   Dispholidus sp. (Boomslang) David & Ineich 1999 [as D. typus] Loveridge 1957 [as D. typus] Pitman 1974 [as D. t. typus] Spawls & Branch 1995 [as D. typus] Spawls, Howell, Hinkel & Menegon 2018 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Grayia tholloni Nandi Water Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Hapsidophrys smaragdinus Common Emerald Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Meizodon (African Smooth Snakes)   Meizodon regularis East African Crowned Smooth Snake Chippaux & Jackson 2019 Schatti 1985 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Hughes 1983 Loveridge 1957 [as P. i. irregularis] Pitman 1974 [as P. i. irregularis] Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Philothamnus battersbyi Northeastern Green Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Philothamnus bequaerti Uganda Green Snake Largen & Rasmussen 1993 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Philothamnus carinatus Barred Green Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Philothamnus heterodermus Emerald Green Snake Hughes 1983 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Philothamnus heterolepidotus Slender Green Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Philothamnus hoplogaster Southeastern Green Snake Spawls, Howell, Drewes & Ashe 2002   Philothamnus hughesi Congolese Green Snake Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Philothamnus nitidus Cameroon Green Snake Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Philothamnus ruandae Rwanda Emerald Green Snake Hughes 1983 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Philothamnus semivariegatus Variegated Bush Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps florulentus Egyptian Whip Snake Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Broadley & Wallach 2002 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Scaphiophis raffreyi Ethiopian Hook-nosed Snake Broadley 1994 Spawls, Howell, Drewes & Ashe 2002 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus obtusus Egyptian Cat Snake Spawls, Howell, Drewes & Ashe 2002 [as dhara] Spawls, Howell, Hinkel & Menegon 2018 [as dhara] Spawls, Malonza & Beraduccii 2021 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Broadley & Wallach 2002 Broadley 2001 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Thrasops (Black Tree Snakes)   Thrasops jacksonii Common Black Tree Snake Broadley & Wallach 2002 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas adamanteus Eastern Powdered Cat Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Hughes 1983 [as pulverulenta] Loveridge 1957 [as pulverulenta] Pitman 1974 [as pulverulenta] Spawls, Howell, Drewes & Ashe 2002 [as pulverulenta] Spawls, Howell, Hinkel & Menegon 2018 [as pulverulenta] Vonesh 1998 [as pulverulenta]   Toxicodryas vexator Eastern Black-and-Yellow Tree Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Hughes 1983 [as blandingii] Loveridge 1957 [as blandingii] Pitman 1974 [as blandingii] Spawls & Branch 1995 [as blandingii] Spawls, Howell, Drewes & Ashe 2002 [as blandingii] Spawls, Howell, Hinkel & Menegon 2018 [as blandingii] Vonesh 1998 [as blandingii]  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba David & Ineich 1999 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Dendroaspis polylepis Black Mamba David & Ineich 1999 Haakansson & Madsen 1983 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Elapsoidea (African Garter Snakes)   Elapsoidea laticincta Sudanese Garter Snake David & Ineich 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Elapsoidea loveridgei Northeastern Garter Snake Broadley 1971 David & Ineich 1999 Jakobsen 1997 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Elapsoidea semiannulata Half-banded Garter Snake Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja ashei East African Spitting Cobra Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014 Wuster & Broadley 2007   Naja haje Egyptian Cobra Pitman 1974 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Trape, Chirio, Broadley & Wuster 2009   Naja nigricollis Black-necked Spitting Cobra Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014 Wuster & Broadley 2007   Naja subfulva Brown Forest Cobra Behangana 2004 [as N. melanoleuca] David & Ineich 1999 [as N. melanoleuca] Hughes 1983 [as N. melanoleuca] Loveridge 1957 [as N. melanoleuca] Pitman 1974 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Spawls, Howell, Drewes & Ashe 2002 [as N. melanoleuca] Spawls, Howell, Hinkel & Menegon 2018 [as N. melanoleuca] Vonesh 1998 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018 Genus Pseudohaje (Tree Cobras)   Pseudohaje goldii Central African Tree Cobra David & Ineich 1999 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus lunulatus Reticulated Centipede Eater Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Aparallactus modestus Western Forest Centipede Eater Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake David & Ineich 1999 Laurent 1950 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake David & Ineich 1999 Hughes 1983 Largen & Rasmussen 1993 Laurent 1950 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Hughes 1997 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Boaedon lineatus Lined House Snake Vonesh 1998   Boaedon olivaceus Olive House Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Boaedon radfordi Lendu Plateau House Snake Greenbaum, Portillo, Jackson & Kusamba 2015 Spawls, Howell, Hinkel & Menegon 2018 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus lineatus Red-and-Black Striped Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Buhoma (African Forest Snakes)   Buhoma depressiceps Pale-headed Forest Snake Chippaux & Jackson 2019 Hughes 1983 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Hughes 2004 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Duberria (African Slug Eaters)   Duberria shirana Shire Slug Eater Broadley & Howell 1991 [as D. lutrix atriventris] Pitman 1974 [as D. lutrix atriventris] Spawls, Howell, Drewes & Ashe 2002 [as D. lutrix] Spawls, Howell, Hinkel & Menegon 2018 [as D. lutrix] Vonesh 1998 [as D. lutrix atriventris] Genus Gonionotophis (Lesser File Snakes)   Gonionotophis brussauxi Eastern Forest Dwarf File Snake Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis kelleri Stripe-bellied Bark Snake Spawls, Howell, Hinkel & Menegon 2018 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Kladirostratus (Striped Beaked Snakes)   Kladirostratus togoensis Western Striped Beaked Snake Chirio & Ineich 1991 Keates, Conradie, Greenbaum & Edwards 2019 Pitman 1974 Genus Limaformosa (Venus File Snakes)   Limaformosa chanleri Unicolor File Snake Spawls, Howell, Hinkel & Menegon 2018   Limaformosa savorgnani Central African Forest File Snake Lanza & Broadley 2014 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Lycophidion (African Wolf Snakes)   Lycophidion depressirostre Flat-snouted Wolf Snake Broadley & Hughes 1993 Broadley 1996 Laurent 1968 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Lycophidion jacksoni Kilimanjaro Wolf Snake Broadley & Howell 1991 Broadley & Hughes 1993 [as L. capense] Broadley 1996 Laurent 1968 Pitman 1974 [as L. c. capense] Spawls, Howell, Drewes & Ashe 2002 [as L. capense] Spawls, Howell, Hinkel & Menegon 2018 [as L. capense] Trape 2021   Lycophidion laterale Masked Wolf Snake Broadley 1996 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Lycophidion ornatum Ornate Wolf Snake Broadley & Hughes 1993 Broadley 1996 Hughes 1983 Laurent 1968 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Mehelya stenophthalmus Small-eyed File Snake Hughes 1983 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Polemon (African Snake Eaters)   Polemon christyi Eastern Snake Eater Hughes 1983 Loveridge 1957 Pitman 1974 Portillo, Branch, Tilbury, Nagy, Hughes, Kusamba, Muninga, Aristote, Behangana & Greenbaum 2019 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Polemon collaris Fawn-headed Snake Eater Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Polemon fulvicollis Yellow-necked Snake Eater Hughes 1983 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Polemon gabonensis Gabon Snake Eater Hughes 1983 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Polemon graueri Central Lakes Snake Eater Loveridge 1957 Pitman 1974 Portillo, Branch, Tilbury, Nagy, Hughes, Kusamba, Muninga, Aristote, Behangana & Greenbaum 2019 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Prosymna (Shovel-snout Snakes)   Prosymna ambigua Central African Shovel-snout Broadley 1980 Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis mossambicus East African Olive Grass Snake Hughes 1983 [as P. phillipsi] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Vonesh 1998 [as P. phillipsii]   Psammophis rukwae Lake Rukwa Sand Snake Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis sudanensis Sudan Sand Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis tanganicus Tanganyika Sand Snake Broadley & Howell 1991 Loveridge 1957 Pitman 1974   Psammophis trivirgatus Southern Speckled Sand Snake Lanza 1990 Largen & Rasmussen 1993 Pitman 1974 Sindaco, Venchi & Grieco 2013 [as P. punctulatus] Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Loveridge 1957 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002   Rhamphiophis rostratus Rufous Beaked Snake Chirio & Ineich 1991  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops emini Karagwe Thread Snake Broadley & Wallach 2007 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls, Howell, Hinkel & Menegon 2018   Leptotyphlops pitmani Uganda Thread Snake Broadley & Wallach 2007 Loveridge 1957 [as L. conjunctus] McDiarmid, Campbell & Toure 1999 [as L. conjunctus] Pitman 1974 [as L. conjunctus] Spawls, Howell, Hinkel & Menegon 2018  Family Micrelapidae (Short-jawed Rear-fanged Snakes) Genus Micrelaps (Afro-Asian Black-headed Snakes)   Micrelaps vaillanti Somali Black-headed Snake Lanza 1990 [as M. boettgeri] Rasmussen 2002 Spawls, Howell, Drewes & Ashe 2002 [as M. boettgeri] Spawls, Howell, Hinkel & Menegon 2018  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Chippaux & Jackson 2019 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Stimson 1969   Python sebae Northern Rock Python Behangana 2004 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops angolensis Golden Blind Snake Broadley & Wallach 2009 Hahn 1980 Hughes 1983 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wallach, Williams & Boundy 2014   Afrotyphlops brevis Somali Giant Blind Snake Broadley & Wallach 2009 Hahn 1980   Afrotyphlops congestus Marbled Blind Snake Broadley & Wallach 2009 Hahn 1980 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wallach, Williams & Boundy 2014   Afrotyphlops lineolatus Lineolate Blind Snake Broadley & Wallach 2009 Hahn 1980 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Broadley & Wallach 2009 Hahn 1980 Hughes 1983 Pitman 1974 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014 Genus Letheobia (African Gracile Blind Snakes)   Letheobia graueri Lake Tanganyika Gracile Blind Snake Broadley & Wallach 2007 Spawls, Howell, Hinkel & Menegon 2018  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   * Atheris acuminata Acuminate Bush Viper Broadley 1998 David & Ineich 1999 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Atheris hispida Rough-scaled Bush Viper Broadley 1998 David & Ineich 1999 Groen, Bok, Tiemann & Verspui 2019 Hughes 1983 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Wallach, Williams & Boundy 2014   Atheris nitschei Great Lakes Bush Viper Broadley 1998 David & Ineich 1999 Hughes 1983 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wallach, Williams & Boundy 2014   Atheris squamigera Southern Green Bush Viper David & Ineich 1999 Hughes 1983 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder David & Ineich 1999 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Bitis gabonica Gaboon Adder Broadley 1983 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Bitis nasicornis African Nose-horned Viper David & Ineich 1999 Hughes 1983 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998 Genus Causus (Night Adders)   Causus lichtensteinii Forest Night Adder David & Ineich 1999 Hughes 1983 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018 Vonesh 1998   Causus maculatus West African Night Adder David & Ineich 1999 Hughes 1983 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Causus resimus Green Night Adder David & Ineich 1999 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Causus rhombeatus Rhombic Night Adder David & Ineich 1999 Loveridge 1957 Pitman 1974 Spawls & Branch 1995 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa neumanni Kenyan Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelomedusa schweinfurthi Central African Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios chapini Central African Mud Turtle Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Pelusios gabonensis Forest Mud Turtle Czurda, Jackson, Muhanguzi & Doherty-Bone 2020 Iverson 1992 King & Burke 1989 Spawls, Howell, Hinkel & Menegon 2018   Pelusios rhodesianus Zambian Mud Turtle Broadley & Boycott 2008 Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Pelusios williamsi Lake Victoria Mud Turtle Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys belliana Central African Hinge-back Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Hinkel & Menegon 2018   Kinixys erosa Forest Hinge-back Tortoise Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Luiselli & Diagne 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018   Kinixys nogueyi Western Savanna Hinge-back Tortoise Fritz & Havas 2006 [as belliana] Iverson 1992 [as belliana] King & Burke 1989 [as belliana] McCord, Joseph-Ouni & Tabaka 2005 [as belliana] Spawls, Howell, Drewes & Ashe 2002 [as belliana] Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018  Family Trionychidae (Softshell Turtles) Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Spawls, Howell, Drewes & Ashe 2002 Spawls, Howell, Hinkel & Menegon 2018