Reptiles of Langkawi  [Malaysia: West Malaysia: Langkawi Archipelago] Related checklists:  Malaysia - West Malaysia - Langkawi Archipelago Bibliography of the reptiles of Langkawi Taxonomic history of the reptiles of Langkawi Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura crucigera Northern Masked Pricklenape Grismer 2008 Grismer 2011 [as "A. cf. crucigera"] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Aphaniotis (Earless Agamas)   Aphaniotis fusca Blue-mouthed Earless Agama Grismer 2011 Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Bronchocela rayaensis Langkawi Long-tailed Agama Grismer & Quah 2019 Grismer, Wood, Lee, Quah, Anuar, Ngadi & Sites 2015 Zug, Mulcahy & Vindum 2017 Genus Calotes (South Asian Forest Agamas)   Calotes emma Forest Crested Agama Grismer 2008 Grismer 2011 Genus Draco (Gliding Dragons)   Draco blanfordii Tavoy Gliding Dragon Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006   Draco maculatus Black-spotted Gliding Dragon Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Leiolepis (Butterfly Agamas)   Leiolepis belliana Common Butterfly Agama Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   * Cnemaspis mahsuriae Mount Raya Rock Gecko Grismer & Quah 2019 Grismer, Wood, Quah, Anuar, Ngadi & Ahmad 2015 Quah, Wood, Anuar, Muin & Grismer 2020   Cnemaspis monachorum Wanaram Temple Rock Gecko Grismer 2011 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Quah, Wood, Anuar, Muin & Grismer 2020   * Cnemaspis roticanai Roti Canai Rock Gecko Grismer & Chan 2010 Grismer 2011 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Quah, Wood, Anuar, Muin & Grismer 2020 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus brevipalmatus Short-handed Bent-toed Gecko Grismer & Quah 2019 Grismer, Wood, Quah, Anuar, Ngadi & Ahmad 2015   * Cyrtodactylus langkawiensis Langkawi Island Bent-toed Gecko Grismer & Quah 2019 Grismer, Wood, Anuar, Quah, Muin, Mohamed, Chan, Sumarli, Loredo & Heinz 2014 Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012   Cyrtodactylus macrotuberculatus Large-tubercled Bent-toed Gecko Grismer 2011 Grismer, Wood, Anuar, Quah, Muin, Mohamed, Chan, Sumarli, Loredo & Heinz 2014 Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012   Cyrtodactylus quadrivirgatus Four-striped Bent-toed Gecko Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Gekko (Tokays)   Gekko cicakterbang Malaysian Gliding Gecko Grismer 2011 [as Ptychozoon lionotum]   Gekko gecko Tokay Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Gekko kuhli Wide-lobed Gliding Gecko Grismer 2008 [as Ptychozoon kuhli] Grismer 2011 [as Ptychozoon kuhli] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 [as Ptychozoon kuhli] Lim, Wira, Chan, Daicus & Norhayati 2010 [as Ptychozoon kuhli]   Gekko monarchus Spotted House Gecko Grismer 2008 Grismer 2011 Lim, Wira, Chan, Daicus & Norhayati 2010   Gekko smithii Greater Green-eyed Gecko Grismer 2008 Grismer 2011 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus craspedotus Malayan Frilled Gecko Grismer 2011   Hemidactylus frenatus Asian House Gecko Grismer 2008 Grismer 2011 [by implication] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Hemidactylus platyurus Flat-tailed Gecko Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Scincidae (Skinks) Genus Dasia (Indo-Malayan Tree Skinks)   Dasia olivacea Olive Tree Skink Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Grismer 2008 Grismer 2011 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Eutropis (South Asian Sun Skinks)   Eutropis macularia Speckled Forest Skink Grismer 2008 Grismer 2011 Lim, Wira, Chan, Daicus & Norhayati 2010   Eutropis multifasciata Many-lined Sun Skink Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Scincella (South Asian and American Ground Skinks)   Scincella melanosticta Black-spotted Ground Skink Grismer & Quah 2019 Grismer, Wood, Lee, Quah, Anuar, Ngadi & Sites 2015 Grismer, Wood, Quah, Anuar, Ngadi & Ahmad 2015 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Toenayar (Nine-keeled Sun Skink)   Toenayar novemcarinata Nine-keeled Sun Skink Grismer & Quah 2019 Grismer, Wood, Quah, Anuar, Ngadi & Ahmad 2015 Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   Tytthoscincus butleri Bintang Leaf-litter Skink Grismer & Quah 2019 Grismer 2008 [as langkawiensis] Grismer 2011 [as langkawiensis] Grismer, Wood, Quah, Anuar, Ngadi, Mohd-Izam & Ahmad 2017 Lim, Wira, Chan, Daicus & Norhayati 2010 [as Sphenomorphus langkawiensis]  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus nebulosus Clouded Monitor Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Varanus salvator Asian Water Monitor Grismer 2008 Grismer 2011 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus javanicus Java File Snake Grismer 2008 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla fasciolata Speckle-headed Whip Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Zimmerer 2004   Ahaetulla nasuta Long-nosed Green Whip Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Zimmerer 2004   Ahaetulla prasina Short-nosed Whip Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Boiga (Australasian Cat Snakes)   Boiga cyanea Green Cat Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Boiga cynodon Dog-toothed Cat Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Boiga melanota Western Mangrove Cat Snake Grismer 2008 [as B. dendrophila] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 [as B. dendrophila] Lim, Wira, Chan, Daicus & Norhayati 2010 [as B. dendrophila] Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Calamaria schlegeli Pink-headed Reed Snake Grismer 2008 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Chrysopelea paradisi Paradise Flying Tree Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus flavolineatus Yellow-striped Rat Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis caudolineatus Malayan Stripe-tailed Bronzeback Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Dendrelaphis formosus Elegant Bronzeback Grismer 2008 Lim, Wira, Chan, Daicus & Norhayati 2010   Dendrelaphis kopsteini Red-necked Bronzeback Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 [as cyanochloris] Lim, Wira, Chan, Daicus & Norhayati 2010   Dendrelaphis pictus Southern Painted Bronzeback Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops rubescens Keel-bellied Vine Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Liopeltis (Ringneck Snakes)   Liopeltis tricolor Malayan Ringneck Snake Lalremsanga, Bal, Vogel & Biakzuala 2022 Som, Grismer, Wood, Quah, Brown, Diesmos, Weinell & Stuart 2020 Genus Lycodon (Oriental Wolf Snakes)   Lycodon subannulatus Malayan Bridle Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Macrocalamus (Peninsular Mountain Reed Snakes)   Macrocalamus lateralis Striped Mountain Reed Snake Quah, Anuar, Grismer, Wood & Nor 2019 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas carinata Malayan Keeled Rat Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis collaris Collared Black-headed Snake Issem 2008 Genus Xenochrophis (Oriental Painted Keelbacks)   Xenochrophis trianguligerus South Asian Triangle Keelback Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis ruffus Red-tailed Pipe Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus candidus Malayan Krait Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Calliophis (South Asian Coral Snakes)   Calliophis maculiceps Small-spotted Coral Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Naja (Typical Cobras)   Naja kaouthia Monocellate Cobra Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Homalopsidae (Australasian Mud Snakes) Genus Enhydris (Asian Mud Snakes)   Enhydris enhydris Striped Mud Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Homalopsis (Masked Water Snakes)   Homalopsis buccata Malayan Masked Water Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Subsessor (Southeast Asian Mud Snake)   Subsessor bocourti Southeast Asian Mud Snake Grismer 2008 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas margaritophorus White-spotted Slug Eater Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Typhlopidae (Typical Blind Snakes) Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Viperidae (Vipers) Genus Calloselasma (Malayan Pitviper)   Calloselasma rhodostoma Malayan Pitviper Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis convictus Indo-Malayan Mountain Pitviper Grismer 2008 [as monticola] Gumprecht & Issem 2013 Lim, Wira, Chan, Daicus & Norhayati 2010 [as monticola] Genus Popeia (South Asian Bamboo Pitvipers)   Popeia sabahi Sunda Bamboo Pitviper Grismer 2008 [as fucata] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 [as fucata] Lim, Wira, Chan, Daicus & Norhayati 2010 [as fucata] Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus purpureomaculatus Mangrove Pitviper Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010   Trimeresurus venustus Venus Pitviper Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus wagleri Speckled Pitviper Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis unicolor Common Asian Sunbeam Snake Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Heosemys (Forest Turtles)   Heosemys annandalii Yellow-headed Temple Turtle Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010 Genus Siebenrockiella (Black Marsh Turtle and Philippine Pond Turtle)   Siebenrockiella crassicollis Black Marsh Turtle Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010  Family Trionychidae (Softshell Turtles) Genus Amyda (Southeast Asian Softshell Turtles)   Amyda ornata Black-rayed Softshell Turtle Grismer 2008 [as A. cartilaginea] Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 [as A. cartilaginea] Lim, Wira, Chan, Daicus & Norhayati 2010 [as A. cartilaginea] Genus Dogania (Malayan Softshell Turtle)   Dogania subplana Malayan Softshell Turtle Grismer 2008 Grismer, Youmans, Wood, Ponce, Wright, Jones, Johnson, Sanders, Gower, Yaakob & Lim 2006 Lim, Wira, Chan, Daicus & Norhayati 2010