Reptiles of Namibia    Bibliography of the reptiles of Namibia Taxonomic history of the reptiles of Namibia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Dalophia (Blunt-tailed Worm Lizards)   Dalophia colobura Sesheke Blunt-tailed Worm Lizard Gans 2005 Mertens 1955   Dalophia longicauda Long-tailed Worm Lizard Branch 1998 Broadley, Gans & Visser 1976 Griffin 2003 Herrmann & Branch 2013   Dalophia pistillum Zambezi Blunt-tailed Worm Lizard Broadley, Gans & Visser 1976 Griffin 2003 Herrmann & Branch 2013 Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis anchietae Mossamedes Spade-snouted Worm Lizard Bauer, Branch & Haacke 1993 Branch 1998 Broadley, Gans & Visser 1976 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Monopeltis infuscata Dusky Spade-snouted Worm Lizard Branch 1998 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group B"] Gans 2005 Griffin 2003 Herrmann & Branch 2013   Monopeltis leonhardi Kalahari Spade-snouted Worm Lizard Branch 1998 Broadley, Gans & Visser 1976 Griffin 2003 Herrmann & Branch 2013   Monopeltis mauricei Botswana Spade-snouted Worm Lizard Griffin 2003 Herrmann & Branch 2013 Genus Zygaspis (Purple Round-headed Worm Lizards)   Zygaspis nigra Black Round-headed Worm Lizard Branch 1998 Broadley & Broadley 1997 Gans 2005 Griffin 2003 Herrmann & Branch 2013   Zygaspis quadrifrons Kalahari Round-headed Worm Lizard Broadley & Broadley 1997 Gans 2005 Griffin 2003 Herrmann & Branch 2013 Mertens 1955

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Cunningham, Maartens & Prickett 2018 Fuchs, Mertens & Wermuth 1974 Griffin 2003 Herrmann & Branch 2013 King & Burke 1989 Mertens 1955 Mertens 1971 Wermuth & Mertens 1977

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus margaritae Kunene Ridgeback Agama Griffin 2003 [as atricollis] Herrmann & Branch 2013 [as A. cyanocephalus] Mertens 1955 [as atricollis] Mertens 1971 [as atricollis] Wagner, Butler, Ceriaco & Bauer 2021 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018 [as A. cyanocephalus] Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Agama anchietae Western Rock Agama Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Agama armata Zambezi Spiny Agama Griffin 2003 Herrmann & Branch 2013   Agama atra Southern Rock Agama Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   * Agama etoshae Etosha Agama Branch 1998 Griffin 2003 Herrmann & Branch 2013   Agama hispida Southern Spiny Agama Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Agama knobeli Namibian Black Agama Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Wermuth 1967   Agama planiceps Damara Rock Agama Bauer, Branch & Haacke 1993 Boehme, Wagner, Malonza, Loetters & Koehler 2005 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Bauer, Branch & Haacke 1993 Branch 1998 Glaw 2015 Griffin 2003 Herrmann & Branch 2013 Klaver & Boehme 1997 Mertens 1955 Tilbury 2010   Chamaeleo namaquensis Namaqua Chameleon Bauer, Branch & Haacke 1993 Branch 1998 Glaw 2015 Griffin 2003 Herrmann & Branch 2013 Klaver & Boehme 1997 Mertens 1955 Mertens 1971 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Karusasaurus (Karoo Girdled Lizards)   * Karusasaurus jordani Northern Karoo Girdled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Karusasaurus polyzonus Southern Karoo Girdled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Namazonurus (Namaqua Girdled Lizards)   * Namazonurus campbelli Helmeringhausen Girdled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Namazonurus namaquensis Karasburg Girdled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Namazonurus pustulatus Herero Girdled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Platysaurus (Flat Lizards)   Platysaurus attenboroughi Karasburg Flat Lizard Branch 1998 [as P. capensis] Broadley 1978 [as P. capensis] Griffin 2003 [as P. capensis] Herrmann & Branch 2013 [as P. capensis] Mertens 1955 [as P. capensis] Mertens 1971 [as P. capensis] Whiting, Branch, Pepper & Keogh 2015  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   * Afroedura africana Southwest African Flat Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Jacobsen, Kuhn, Jackman & Bauer 2014 Mertens 1955   * Afroedura otjihipa Otjihipa Flat Gecko Conradie, Schmitz, Lobon-Rovira, Becker, Vaz Pinto & Hauptfleisch 2022   * Afroedura tirasensis Luderitz Flat Gecko Branch 1998 Griffin 2003 Haacke 1965 Jacobsen, Kuhn, Jackman & Bauer 2014 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus angulifer Giant Ground Gecko Branch 1998 Griffin 2003 Heinz, Brennan, Jackman & Bauer 2021 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Chondrodactylus bibronii Tubercled Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Heinz, Brennan, Jackman & Bauer 2021 Herrmann & Branch 2013   Chondrodactylus fitzsimonsi Kunene Thick-toed Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Heinz, Brennan, Jackman & Bauer 2021 Herrmann & Branch 2013 Mertens 1955   Chondrodactylus laevigatus Karas Thick-toed Gecko Griffin 2003 Heinz, Brennan, Jackman & Bauer 2021 Herrmann & Branch 2013 Marques, Ceriaco, Blackburn & Bauer 2018 Mertens 1955 Mertens 1971   Chondrodactylus pulitzerae Pico Azevedo Thick-toed Gecko Ceriaco, Bauer, Blackburn & Lavres 2014 Griffin 2003 Heinz, Brennan, Jackman & Bauer 2021 Mertens 1955 Genus Goggia (South African Dwarf Leaf-toed Geckos)   Goggia gemmula Richtersveld Dwarf Leaf-toed Gecko Bauer, Good & Branch 1997 Branch 1998 Griffin 2003 Heinicke, Turk & Bauer 2017 Herrmann & Branch 2013   Goggia lineata Striped Dwarf Leaf-toed Gecko Bauer, Good & Branch 1997 Branch 1998 Branch, Bauer & Good 1995 Griffin 2003 Heinicke, Turk & Bauer 2017 Herrmann & Branch 2013 Mertens 1955 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus benguellensis Benguela Half-toed Gecko Branch 1998 [as longicephalus] Ceriaco, Agarwal, Marques & Bauer 2020 Griffin 2003 [as longicephalus] Herrmann & Branch 2013 [as longicephalus]   Hemidactylus mabouia Cosmopolitan House Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus bradfieldi Namibian Dwarf Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Lygodactylus capensis Cape Dwarf Gecko Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Lygodactylus chobiensis Chobe Dwarf Gecko Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Lygodactylus lawrencei Kaokoveld Dwarf Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Lygodactylus nyaneka Nyaneka Dwarf Gecko Marques, Ceriaco, Buehler, Bandeira, Janota & Bauer 2020 Genus Narudasia (Festive Gecko)   * Narudasia festiva Festive Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Pachydactylus (Typical Thick-toed Geckos)   * Pachydactylus acuminatus Namibian Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013 Mertens 1955   Pachydactylus atorquatus Augrabies Thick-toed Gecko Bauer, Barts & Hulbert 2006 Herrmann & Branch 2013   Pachydactylus barnardi Garies Thick-toed Gecko Bauer, Ceriaco, Heinicke & Blackburn 2015 Herrmann & Branch 2013   * Pachydactylus bicolor Velvety Thick-toed Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Pachydactylus boehmei Uisib Thick-toed Gecko Bauer 2010 Herrmann & Branch 2013   Pachydactylus capensis Common Cape Thick-toed Gecko Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Pachydactylus caraculicus Angolan Banded Thick-toed Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Pachydactylus carinatus Richtersveld Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   * Pachydactylus etultra Sossus Gecko Bauer & Lamb 2002 Branch, Bauer, Jackman & Heinicke 2011 Herrmann & Branch 2013   * Pachydactylus fasciatus Damaraland Banded Thick-toed Gecko Bauer, Branch & Haacke 1993 Bauer, Lamb & Branch 2006 Herrmann & Branch 2013 Huber, Barts & Schneider 2011 Mertens 1955   * Pachydactylus gaiasensis Brandberg Thick-toed Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   * Pachydactylus griffini Keetmanshoop Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus haackei Great Namaqualand Thick-toed Gecko Branch 1998 Branch, Bauer & Good 1996 Griffin 2003 Herrmann & Branch 2013   * Pachydactylus kobosensis Kobos Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013 Mertens 1955   * Pachydactylus kochii Namib Ghost Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Pachydactylus latirostris Broad-nosed Thick-toed Gecko Bauer, Heinicke, Jackman & Branch 2011 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   * Pachydactylus maraisi Swakopmund Thick-toed Gecko Heinicke, Adderly, Bauer & Jackman 2011 Herrmann & Branch 2013   * Pachydactylus mclachlani Orange River Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus monicae Sendelingsdrif Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus montanus Karas Mountains Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus namaquensis Namaqua Thick-toed Gecko Branch 1998 Branch, Bauer & Good 1996 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Pachydactylus oreophilus Kaokoveld Thick-toed Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   * Pachydactylus otaviensis Otavi Highlands Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   * Pachydactylus parascutatus Sesfontein Thick-toed Gecko Bauer, Lamb & Branch 2002 Griffin 2003 Herrmann & Branch 2013   Pachydactylus punctatus Speckled Thick-toed Gecko Bauer & Branch 1995 Bauer & Lamb 2002 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Pachydactylus purcelli Karoo Spotted Thick-toed Gecko Bauer, Lamb & Branch 2006 Branch 1998 Herrmann & Branch 2013 Mertens 1955   Pachydactylus rangei Namib Web-footed Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Pachydactylus reconditus Windhoek Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   * Pachydactylus robertsi Karasburg Thick-toed Gecko Bauer, Lamb & Branch 2006 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Pachydactylus rugosus Rough Thick-toed Gecko Bauer & Lamb 2002 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Lamb & Bauer 2000 Mertens 1955   * Pachydactylus sansteynae Kuidas Thick-toed Gecko Bauer, Branch & Haacke 1993 Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus scherzi Namib Variable Gecko Bauer & Branch 1995 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Pachydactylus scutatus Armored Thick-toed Gecko Bauer, Branch & Haacke 1993 Bauer, Lamb & Branch 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   * Pachydactylus serval Western Spotted Thick-toed Gecko Bauer, Lamb & Branch 2006 Branch 1998 Herrmann & Branch 2013 Mertens 1955   Pachydactylus vanzyli Kaoko Web-footed Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Pachydactylus visseri Ai-Ais Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus wahlbergii Kalahari Ground Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Pachydactylus waterbergensis Waterberg Thick-toed Gecko Bauer, Jackman, Branch & Marais 2007 Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   Pachydactylus weberi Klipfontein Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013   * Pachydactylus werneri Khan River Thick-toed Gecko Bauer, Lamb & Branch 2006 Herrmann & Branch 2013 Mertens 1955 Genus Ptenopus (African Barking Geckos)   * Ptenopus carpi Banded Barking Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Ptenopus garrulus Common Barking Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Ptenopus kochi Interdune Barking Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Genus Rhoptropus (Southwest African Day Geckos)   Rhoptropus afer Coastal Namib Day Gecko Bauer, Branch & Haacke 1993 Branch 1998 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Rhoptropus barnardi Kunene River Day Gecko Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Rhoptropus biporosus Kaokoveld Day Gecko Branch 1998 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Rhoptropus boultoni Pico Azevedo Day Gecko Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   * Rhoptropus bradfieldi Messum River Day Gecko Bauer & Lamb 2001 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   * Rhoptropus diporus Damara Day Gecko Bauer & Lamb 2001 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013  Family Gerrhosauridae (Plated Lizards) Genus Cordylosaurus (Dwarf Plated Lizard)   Cordylosaurus subtessellatus Dwarf Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus auritus Kalahari Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 [as multilineatus] Mertens 1955 Mertens 1971   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Gerrhosaurus intermedius Eastern Black-lined Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 [as nigrolineatus] Bauer, Branch & Haacke 1993 [as nigrolineatus] Branch 1998 [as nigrolineatus] Griffin 2003 [as nigrolineatus] Herrmann & Branch 2013 [as nigrolineatus] Mertens 1955 [as n. nigrolineatus] Mertens 1971 [as nigrolineatus]   Gerrhosaurus skoogi Desert Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Gerrhosaurus typicus Namaqua Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Herrmann & Branch 2013 Genus Matobosaurus (Giant Plated Lizards)   Matobosaurus maltzahni Western Giant Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 [as validus] Mertens 1955 Wermuth 1968  Family Lacertidae (Typical Lizards) Genus Heliobolus (Bushveld Lizards)   Heliobolus lugubris Southern African Bushveld Lizard Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Marques, Ceriaco, Heinicke, Chehouri, Conradie, Tolley & Bauer 2022 Mertens 1955 Mertens 1971 Szczerbak 1975 Genus Ichnotropis (Rough-scaled Lizards)   Ichnotropis capensis Southern Rough-scaled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Ichnotropis grandiceps Caprivi Rough-scaled Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Genus Meroles (Southwest African Desert Lizards)   Meroles anchietae Shovel-snouted Lizard Branch 1998 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Meroles ctenodactylus Giant Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   Meroles cuneirostris Wedge-snouted Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Meroles knoxii Round-snouted Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   * Meroles micropholidotus Namibian Small-scaled Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   Meroles reticulatus Reticulated Desert Lizard Branch 1998 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   Meroles squamulosus Rough-scaled Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Meroles suborbitalis Spotted Desert Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975 Genus Nucras (Sandveld Lizards)   * Nucras damarana Damara Sandveld Lizard Bauer, Murdoch & Childers 2020 Branch 1998 [as holubi] Griffin 2003 [as holubi] Herrmann & Branch 2013 [as taeniolata] Mertens 1955 [as holubi] Parker 1936   Nucras intertexta Spotted Sandveld Lizard Bauer, Branch & Haacke 1993 Bauer, Murdoch & Childers 2020 Branch 1998 Broadley 1972 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Nucras tessellata Western Sandveld Lizard Bauer, Murdoch & Childers 2020 Branch 1998 Broadley 1972 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Pedioplanis (African Sand Lizards)   Pedioplanis benguellensis Benguela Sand Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013   * Pedioplanis branchi Karibib Sand Lizard Childers, Kirchhof & Bauer 2021   * Pedioplanis breviceps Short-headed Sand Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   * Pedioplanis gaerdesi Kaokoveld Sand Lizard Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   * Pedioplanis husabensis Husab Sand Lizard Berger-Dell'mour & Mayer 1989 Branch 1998 Griffin 2003 Herrmann & Branch 2013   Pedioplanis inornata Plain Sand Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Szczerbak 1975   Pedioplanis laticeps Cape Sand Lizard Griffin 2003 Herrmann & Branch 2013 Kirchhof, Penner, Roedel & Muller 2017 Mertens 1955 Mertens 1971   Pedioplanis lineoocellata Spotted Sand Lizard Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Szczerbak 1975   * Pedioplanis mayeri Omaruru Sand Lizard Childers, Kirchhof & Bauer 2021   Pedioplanis namaquensis Namaqua Sand Lizard Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   Pedioplanis pulchella Venus Sand Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Szczerbak 1975   * Pedioplanis rubens Waterberg Sand Lizard Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975   * Pedioplanis undata Western Sand Lizard Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Szczerbak 1975  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias gariepensis Gariep Blind Legless Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Acontias kgalagadi Kgalagadi Legless Skink Branch 1998 [as Typhlosaurus l. lineatus] Broadley 1968 [as Typhlosaurus lineatus] Conradie & Bourquin 2013 Griffin 2003 [as Typhlosaurus lineatus] Herrmann & Branch 2013 Lamb, Biswas & Bauer 2010 Mertens 1955 [as Typhlosaurus lineatus] Mertens 1971 [as Typhlosaurus lineatus]   Acontias lineatus Striped Dwarf Legless Skink Branch 1998 Broadley & Greer 1969 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Acontias occidentalis Western Black Legless Skink Branch 1998 Broadley & Greer 1969 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Genus Mochlus (African Supple Skinks)   Mochlus sundevallii Common Savanna Supple Skink Bauer, Branch & Haacke 1993 Broadley 1966 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Panaspis (African Snake-eyed Skinks)   Panaspis maculicollus Spotted-neck Snake-eyed Skink Ceriaco, Branch & Bauer 2018 Griffin 2003 Herrmann & Branch 2013 Jacobsen & Broadley 2000   * Panaspis namibiana Namibian Snake-eyed Skink Bauer, Branch & Haacke 1993 [as wahlbergii] Branch 1998 [as wahlbergii] Ceriaco, Branch & Bauer 2018 Herrmann & Branch 2013 [as wahlbergii] Mertens 1955 [as wahlbergii] Mertens 1971 [as wahlbergii] Genus Scelotes (Dwarf Burrowing Skinks)   Scelotes capensis Namibian Dwarf Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Scelotes sexlineatus Striped Dwarf Burrowing Skink Bauer 2019 Genus Sepsina (Savannah Burrowing Skinks)   * Sepsina alberti Namibian Burrowing Skink Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Sepsina angolensis Angolan Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis acutilabris Wedge-snouted Skink Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis binotata Ovambo Tree Skink Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis capensis Cape Skink Broadley 2000 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis chimbana Chimba Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013   Trachylepis damarana Damara Variable Skink Branch 1998 [as varia] Broadley 2000 [as varia] Griffin 2003 [as varia] Herrmann & Branch 2013 [as varia] Mertens 1955 [as varia] Mertens 1971 [as varia] Peters 1870 Portik, Bauer & Jackman 2010 [as varia] Weinell & Bauer 2018   Trachylepis hoeschi Western Rock Skink Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis laevis Angolan Blue-tailed Skink Bauer, Branch & Haacke 1993 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Trachylepis occidentalis Western Three-striped Skink Bauer, Branch & Haacke 1993 Branch 1998 Broadley 2000 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis punctatissima Montane Speckled Skink Griffin 2003 Herrmann & Branch 2013   Trachylepis punctulata Eastern Variegated Skink Branch 1998 Broadley 1975 Broadley 2000 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 [as longiloba triebneri] Portik & Bauer 2012 Portik, Bauer & Jackman 2010   Trachylepis sparsa Karasburg Tree Skink Branch 1998 Broadley 1977 Broadley 2000 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Trachylepis spilogaster Namibian Tree Skink Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1977 Broadley 2000 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis striata Eastern Striped Skink Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Trachylepis sulcata Koppie Skink Bauer, Branch & Haacke 1993 Branch 1998 Broadley 2000 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Portik, Bauer & Jackman 2010 Portik, Bauer & Jackman 2011   Trachylepis variegata Western Variegated Skink Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1975 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 [as l. longiloba] Mertens 1971 Portik & Bauer 2012 Portik, Bauer & Jackman 2010   Trachylepis wahlbergii Mopane Skink Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1977 Broadley 2000 Griffin 2003 Genus Typhlacontias (Western Burrowing Skinks)   * Typhlacontias brevipes Short-legged Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Haacke 1997   Typhlacontias gracilis Zambian Burrowing Skink Herrmann & Branch 2013   Typhlacontias johnsonii Kaokoveld Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Haacke 1997   Typhlacontias punctatissimus Speckled Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Haacke 1997   Typhlacontias rohani Kalahari Burrowing Skink Branch 1998 Griffin 2003 Herrmann & Branch 2013 Haacke 1997 Mertens 1955 [as Typhlacontias ngamiensis] Genus Typhlosaurus (Blind Legless Skinks)   * Typhlosaurus braini Slender Blind Legless Skink Branch 1998 Broadley 1968 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Typhlosaurus meyeri Sperrgebiet Blind Legless Skink Branch 1998 Broadley 1968 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bauer, Branch & Haacke 1993 Bayless 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   Varanus niloticus Nile Monitor Bayless 2002 Griffin 2003 Herrmann & Branch 2013 Lenz in Pianka, King & King (eds.) 2004 Mertens 1971

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Genus Dasypeltis (African Egg Eaters)   Dasypeltis scabra Common African Egg Eater Bates & Broadley 2018 Gans 1959 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Pietersen, Verburght & Davies 2021 Genus Limnophis (Striped Swamp Snakes)   Limnophis bangweolicus Eastern Striped Swamp Snake Conradie, Deepak, Keates & Gower 2020 Griffin 2003 Herrmann & Branch 2013 Genus Mopanveldophis (Cunene Whip Snake)   * Mopanveldophis zebrinus Cunene Whip Snake Bauer, Lamb, Branch & Babb 2001 Branch 1998 Broadley & Schatti 1997 Broadley & Schatti 2000 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Broadley 1966 Griffin 2003 Herrmann & Branch 2013 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 [as P. i. irregularis] Mertens 1971 [as P. irregularis]   Philothamnus hoplogaster Southeastern Green Snake Griffin 2003 Herrmann & Branch 2013   Philothamnus ornatus Ornate Green Snake Griffin 2003 Herrmann & Branch 2013   Philothamnus semivariegatus Variegated Bush Snake Bauer, Branch & Haacke 1993 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus beetzii Namib Tiger Snake Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Telescopus finkeldeyi Damara Cat Snake Haacke 2013   Telescopus semiannulatus Southern Tiger Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Loveridge 1957 Mertens 1955 Mertens 1971 Genus Thelotornis (African Vine Snakes)   Thelotornis capensis Southern Vine Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Wallach 2002 Broadley 1983 Broadley 2001 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Spawls & Branch 1995  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps lubricus African Coral Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Baldwin 2006 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Wallach, Williams & Boundy 2014   Aspidelaps scutatus Shield-nosed Coral Snake Branch 1998 Broadley & Baldwin 2006 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Wallach, Williams & Boundy 2014 Genus Dendroaspis (Mambas)   Dendroaspis polylepis Black Mamba Bauer, Branch & Haacke 1993 Broadley 1983 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Haakansson & Madsen 1983 Mertens 1955 Mertens 1971 Spawls & Branch 1995 Genus Elapsoidea (African Garter Snakes)   Elapsoidea boulengeri Zambezi Garter Snake Broadley 1971 Broadley 1998 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013   Elapsoidea semiannulata Half-banded Garter Snake Branch 1998 Broadley 1983 Broadley 1998 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Spawls & Branch 1995   Elapsoidea sundevallii Southern African Garter Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1971 Broadley 1983 Broadley 1998 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja anchietae Southwest African Cobra Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Wuster 2004 Broadley 1968 Broadley 1983 Broadley 1995 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995   Naja crawshayi Katanga Spitting Cobra Branch 1998 [as N. n. nigricollis] Griffin 2003 [as N. n. nigricollis] Herrmann & Branch 2013 [as N. n. nigricollis]   Naja mossambica Mozambique Spitting Cobra Branch 1998 Broadley 1983 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja nigricincta Western Barred Spitting Cobra Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1968 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja nivea Yellow Cobra Branch 1998 Broadley & Wuster 2004 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis platurus Yellow-bellied Sea Snake Branch 1998 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013  Family Lamprophiidae (House Snakes, etc.) Genus Alopecion (Spotted House Snake)   Alopecion guttatum Spotted House Snake Branch 1998 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Bauer, Branch & Haacke 1993 Broadley 1971 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Amblyodipsas mechowii Elongate Quill-snouted Snake Branch 1998 Broadley 1971 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Amblyodipsas polylepis Common Purple-glossed Snake Broadley 1971 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Wallach, Williams & Boundy 2014   Amblyodipsas ventrimaculata Kalahari Purple-glossed Snake Branch 1998 Broadley 1971 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Genus Aparallactus (African Centipede Eaters)   Aparallactus capensis Cape Centipede Eater Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Bauer, Branch & Haacke 1993 Broadley 1983 Broadley 1991 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995 Wallach, Williams & Boundy 2014   Atractaspis congica Congo Stiletto Snake Broadley 1983 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Spawls & Branch 1995   Atractaspis duerdeni Beaked Stiletto Snake Branch 1998 Broadley 1991 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Herrmann & Branch 2013   Boaedon fradei Moxico House Snake Hallermann, Ceriaco, Schmitz, Ernst, Conradie, Verburgt, Marques & Bauer 2020   Boaedon mentalis Namibe House Snake Hallermann, Ceriaco, Schmitz, Ernst, Conradie, Verburgt, Marques & Bauer 2020 Mertens 1955   Boaedon variegatus Variegated House Snake Hallermann, Ceriaco, Schmitz, Ernst, Conradie, Verburgt, Marques & Bauer 2020 Genus Dipsina (Dwarf Beaked Snake)   Dipsina multimaculata Dwarf Beaked Snake Branch 1998 Broadley 1983 Chirio & Ineich 1991 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Genus Gracililima (Black File Snake)   Gracililima nyassae Black File Snake Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Lanza & Broadley 2014 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Hughes 2000 Griffin 2003 Herrmann & Branch 2013   Hemirhagerrhis viperina Viperine Bark Snake Bauer, Branch & Haacke 1993 Bauer, Jackman, Branch & Marais 2007 Broadley & Hughes 2000 Broadley 1983 Broadley 1997 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Genus Limaformosa (Venus File Snakes)   Limaformosa capensis Cape File Snake Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013   Limaformosa vernayi Angolan File Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Bauer, Branch & Haacke 1993 Branch 1976 Branch 1998 Broadley 1991 Broadley 1996 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Lycophidion hellmichi Kaokoveld Wolf Snake Branch 1998 Broadley 1991 Broadley 1996 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Lycophidion multimaculatum Many-spotted Wolf Snake Branch 1998 Broadley & Howell 1991 Broadley 1983 Broadley 1991 Broadley 1996 Griffin 2003 Herrmann & Branch 2013 Laurent 1968   Lycophidion namibianum Namibian Wolf Snake Branch 1998 Broadley 1991 Broadley 1996 Griffin 2003 Herrmann & Branch 2013 Genus Prosymna (Shovel-snout Snakes)   Prosymna angolensis Angolan Shovel-snout Branch 1998 Broadley 1980 Broadley 1983 Conradie, Keates, Baptista & Lobon-Rovira 2022 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Prosymna bivittata Two-striped Shovel-snout Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1980 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Prosymna frontalis Southwestern Shovel-snout Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1980 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Prosymna lisima Kalahari Shovel-snout Conradie, Keates, Baptista & Lobon-Rovira 2022   Prosymna visseri Kaokoveld Shovel-snout Bauer, Branch & Haacke 1993 Bauer, Lamb, Branch & Babb 2001 Branch 1998 Griffin 2003 Herrmann & Branch 2013 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis angolensis Dwarf Sand Snake Branch 1998 Broadley 1983 Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1971   Psammophis brevirostris Short-snouted Grass Snake Broadley 2002 Griffin 2003 Herrmann & Branch 2013   Psammophis jallae Zimbabwe Sand Snake Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Psammophis leopardinus Leopard Sand Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002 Griffin 2003 Herrmann & Branch 2013   Psammophis mossambicus East African Olive Grass Snake Broadley & Howell 1991 [as P. phillipsi] Broadley 1977 [as P. phillipsi] Broadley 1983 [as P. phillipsii] Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis namibensis Namib Sand Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Broadley 2002 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 [as P. leightoni]   Psammophis notostictus Karoo Sand Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Broadley 2002 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Psammophis subtaeniatus Western Stripe-bellied Sand Snake Bauer, Branch & Haacke 1993 Broadley 1977 Broadley 1983 Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Psammophis trigrammus Western Sand Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Psammophis trinasalis Kalahari Sand Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1977 Broadley 2002 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Genus Psammophylax (Skaapstekers)   Psammophylax rhombeatus Spotted Skaapsteker Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1976 Griffin 2003   Psammophylax tritaeniatus Southern Striped Skaapsteker Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1976 Griffin 2003 Herrmann & Branch 2013 Loveridge 1957 Mertens 1955 Mertens 1971   Psammophylax variabilis Grey-bellied Skaapsteker Broadley & Howell 1991 Broadley 1976 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Bauer, Branch & Haacke 1993 Griffin 2003 Herrmann & Branch 2013 Loveridge 1957 Mertens 1955 Mertens 1971 Genus Pythonodipsas (Western Keeled Snake)   Pythonodipsas carinata Western Keeled Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis rostratus Rufous Beaked Snake Branch 1998 Griffin 2003 Herrmann & Branch 2013 [as R. oxyrhynchus]  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops conjunctus Cape Thread Snake Herrmann & Branch 2013   Leptotyphlops scutifrons Zambezi Thread Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley & Watson 1976 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Genus Namibiana (Southwest African Thread Snakes)   Namibiana gracilior Slender Thread Snake Branch 1998 Broadley & Broadley 1999 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999   Namibiana labialis Damara Thread Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Broadley 1999 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1955 Mertens 1971   Namibiana occidentalis Western Namaqua Thread Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Broadley 1999 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1971  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python anchietae Angolan Python Bauer, Branch & Haacke 1993 Branch 1998 Broadley 1983 Chippaux & Jackson 2019 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971   Python natalensis Southern Rock Python Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Howell 1991 Broadley 1983 Broadley 1984 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 [as sebae] Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops schlegelii Variable Giant Blind Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops boylei Kalahari Beaked Blind Snake Branch 1998 Broadley & Wallach 2009 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1955 Mertens 1971   Rhinotyphlops lalandei Cape Blind Snake Branch 1998 Broadley & Wallach 2009 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1955 Mertens 1971   Rhinotyphlops schinzi Black-spotted Beaked Blind Snake Bauer, Branch & Haacke 1993 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Bauer, Branch & Haacke 1993 Branch 1998 Cunningham, Maartens & Prickett 2018 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1955 Mertens 1971 Spawls & Branch 1995   Bitis caudalis Horned Adder Bauer, Branch & Haacke 1993 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Spawls & Branch 1995   Bitis cornuta Many-horned Adder Branch 1998 Branch 1999 Griffin 2003 Herrmann & Branch 2013 Mertens 1955   * Bitis peringueyi Namib Dwarf Sand Adder Branch 1998 Broadley 1983 Chippaux & Jackson 2019 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Mertens 1955 Mertens 1971 Spawls & Branch 1995   Bitis schneideri Namaqua Dwarf Adder Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995   Bitis xeropaga Desert Mountain Adder Branch 1998 Broadley 1983 Griffin 2003 Herrmann & Branch 2013 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus rhombeatus Rhombic Night Adder Branch 1998 David & Ineich 1999 Griffin 2003 Herrmann & Branch 2013 Spawls & Branch 1995

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Herrmann & Branch 2013 Hughes in Bjorndal (ed.) 1982 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Brongersma in Bjorndal (ed.) 1982 Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Hughes in Bjorndal (ed.) 1982 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Griffin 2003 Herrmann & Branch 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Cunningham, Maartens & Prickett 2018 Griffin 2003 Herrmann & Branch 2013 Hughes in Bjorndal (ed.) 1982 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa subrufa Common Helmeted Turtle Boycott & Bourquin 2008 Griffin 2003 Herrmann & Branch 2013 King & Burke 1989 Mertens 1955 Mertens 1971 Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios bechuanicus Okavango Mud Turtle Fritz & Havas 2006 Griffin 2003 Herrmann & Branch 2013 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios rhodesianus Zambian Mud Turtle Broadley & Boycott 2008 Griffin 2003 Herrmann & Branch 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Chersina (Angulate Tortoise)   Chersina angulata Angulate Tortoise Branch 1998 Griffin 2003 Herrmann & Branch 2013 Hofmeyr 2009 King & Burke 1989 Mertens 1955 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chersobius (Five-toed Dwarf Tortoises)   * Chersobius solus Nama Padloper Branch 1998 [as Homopus sp.] Branch 2007 Herrmann & Branch 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Kinixys (Hinge-back Tortoises)   Kinixys spekii Eastern Savanna Hinge-back Tortoise Griffin 2003 Hailey, Broadley, Boycott & Spawls 2021 Herrmann & Branch 2013 Ihlow, Farooq, Gvozdik, Hofmeyr, Conradie, Campbell, Harvey, Verburgt & Fritz 2019 Kindler, Branch, Hofmeyr, Maran, Siroky, Vences, Harvey, Hauswaldt, Schleicher, Stuckas & Fritz 2012 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Psammobates (South African Star Tortoises)   Psammobates oculiferus African Serrated Star Tortoise Bauer, Branch & Haacke 1993 Fritz & Havas 2006 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Psammobates tentorius African Tent Tortoise Branch 1998 Fritz & Havas 2006 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Bauer, Branch & Haacke 1993 Fritz & Havas 2006 Griffin 2003 Herrmann & Branch 2013 Mertens 1955 Mertens 1971 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Branch 1998 Cunningham, Maartens & Prickett 2018 Fritz & Havas 2006 Griffin 2003 Herrmann & Branch 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021