Reptiles of Nigeria    Bibliography of the reptiles of Nigeria Taxonomic history of the reptiles of Nigeria Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Baikia (Nigerian Worm Lizard)   * Baikia africana Nigerian Worm Lizard Gans 1967 Gans 2005 Genus Cynisca (West African Round-headed Worm Lizards)   * Cynisca gansi Okoloma Worm Lizard Gans 1987 Gans 2005   * Cynisca kigomensis Kigom Hills Worm Lizard Gans 1987 Gans 2005   Cynisca leucura White-tailed Worm Lizard Gans 1987 Gans 2005   * Cynisca nigeriensis Takwashara Worm Lizard Gans 1987 Gans 2005

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 Fuchs, Mertens & Wermuth 1974 King & Burke 1989 [as niloticus] Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   Mecistops cataphractus West African Slender-snouted Crocodile King & Burke 1989 Shirley, Carr, Nestler, Vliet & Brochu 2018 Trape, Trape & Chirio 2012 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 King & Burke 1989 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama agama Central African Rainbow Agama Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019   Agama boueti Mali Agama Trape, Trape & Chirio 2012   Agama doriae Nigerian Agama Butler 1986 Chirio & LeBreton 2007 Grandison 1968 Joger 1982 Mediannikov, Trape & Trape 2012 Moody & Boehme 1984 Trape, Trape & Chirio 2012   Agama gracilimembris Lesser Ground Agama Gartshore 1985 Grandison 1968 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Wagner & Bauer 2011   Agama lebretoni Cameroon Rainbow Agama Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012   Agama parafricana Lesser African Forest Agama Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Nneji, Adeola, Yan, Okeyoyin, Oladipo, Olagunju, Omotoso, Oladipo, Iyiola, Auta, Usman, Abdullahi, Peng, Jin, Murphy, Ugwumba & Che 2018   Agama paragama West African Tree Agama Butler 1986 Grandison 1968 Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012   Agama picticauda Ada Foah Agama Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016   Agama sankaranica Sankaranian Agama Grandison 1968 Joger 1979 Mediannikov, Trape & Trape 2012 Papenfuss 1969 Trape, Trape & Chirio 2012 Wermuth 1967  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo africanus Sahel Chameleon Dunger 1967 [as basiliscus] Glaw 2015 Joger 1981 Klaver & Boehme 1997 Papenfuss 1969 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo gracilis Graceful Chameleon Butler 1986 Dunger 1967 Glaw 2015 Klaver & Boehme 1997 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo necasi Togo Chameleon Chirio & LeBreton 2007 [as quilensis] Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Dunger 1967 Glaw 2015 Joger 1981 Tilbury 2010 Trape, Trape & Chirio 2012 Genus Rhampholeon (African Leaf Chameleons)   Rhampholeon spectrum Spectral Leaf Chameleon Dunger 1967 Glaw 2015 Reid 1986 Tilbury 2010 Trape, Trape & Chirio 2012 Wild 1994 Genus Trioceros (Equatorial Chameleons)   Trioceros cristatus Crested Chameleon Dunger 1967 Glaw 2015 Joger 1990 Klaver & Boehme 1997 Reid 1986 Romer 1953 Tilbury 2010 Trape, Trape & Chirio 2012   Trioceros oweni Central African Three-horned Chameleon Dunger 1967 Glaw 2015 Klaver & Boehme 1997 Romer 1953 Tilbury 2010 Trape, Trape & Chirio 2012   Trioceros quadricornis Four-horned Chameleon Boehme 1975 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010 Trape, Trape & Chirio 2012   Trioceros serratus Serrated Chameleon Barej, Ineich, Gvozdik, Lhermitte-Vallarino, Gonwouo, LeBreton, Bott & Schmitz 2010 Glaw 2015 Trape, Trape & Chirio 2012   Trioceros wiedersheimi Northern Peacock Chameleon Barej, Ineich, Gvozdik, Lhermitte-Vallarino, Gonwouo, LeBreton, Bott & Schmitz 2010 Glaw 2015 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Butler 1986 Dunger 1968 Grismer 1988 Loveridge 1947 Papenfuss 1969 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   * Ancylodactylus gigas Giant Rock Gecko Malonza & Bauer 2022 Perret 1986 Trape & Chirio 2013 Trape, Trape & Chirio 2012   Ancylodactylus koehleri Cameroon Mountains Rock Gecko Bauer & LeBreton 2021   * Ancylodactylus petrodroma Ondo Rock Gecko Malonza & Bauer 2022 Perret 1986 Trape, Trape & Chirio 2012   Ancylodactylus spinicollis Spiny-necked Rock Gecko Perret 1986 Segniagbeto 2021 Trape, Trape & Chirio 2012 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus albituberculatus White-tubercled Half-toed Gecko Trape, Trape & Chirio 2012   Hemidactylus angulatus West African House Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Butler 1986 Cansdale 1955 Dunger 1968 Lajmi, Bansal, Giri & Karanth 2019 Loveridge 1947 Papenfuss 1969 Reid 1986 Romer 1953 Trape, Trape & Chirio 2012   Hemidactylus ansorgii Slender Forest Gecko Chirio & LeBreton 2007 Henle & Boehme 2003 Trape, Trape & Chirio 2012   Hemidactylus echinus Spiny Forest Gecko Reid 1986   Hemidactylus eniangii Cross River Half-toed Gecko Wagner, Leache & Fujita 2014   Hemidactylus mabouia Cosmopolitan House Gecko Dunger 1968 Loveridge 1947 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Reid 1986 Trape, Trape & Chirio 2012 Wermuth 1965   Hemidactylus matschiei Adele Half-toed Gecko Dunger 1968 Loveridge 1947 Trape, Trape & Chirio 2012 Wermuth 1965   Hemidactylus muriceus Prickly Half-toed Gecko Dunger 1968 Loveridge 1947 [as intestinalis] Reid 1986 [as intestinalis] Trape, Trape & Chirio 2012   Hemidactylus richardsonii Central African Half-toed Gecko Dunger 1968 Trape, Trape & Chirio 2012 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus conraui Cameroon Dwarf Gecko Butler 1986 Dunger 1968 Reid 1986 Trape, Trape & Chirio 2012   Lygodactylus fischeri Sierra Leone Dwarf Gecko Dunger 1968 Trape, Trape & Chirio 2012   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Dunger 1968 Trape, Trape & Chirio 2012 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus sthenodactylus Elegant Thin-toed Gecko Arnold 1980 Baha el Din 2006 Dunger 1968 Papenfuss 1969 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Tropiocolotes (Afro-Asian Desert Dwarf Geckos)   Tropiocolotes tripolitanus Tripoli Dwarf Gecko Trape, Trape & Chirio 2012  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Arnold 1983 Brito, Rebelo, Crochet & Geniez 2008 Dunger 1967 Papenfuss 1969 Salvador 1982 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Acanthodactylus boueti Benin Fringe-toed Lizard Salvador 1982 Trape, Trape & Chirio 2012   Acanthodactylus guineensis Guinea Fringe-toed Lizard Arnold 1983 Dunger 1967 Joger & Lambert 1996 Meinig & Boehme 2002 Papenfuss 1969 Salvador 1982 Szczerbak 1975 Trape, Trape & Chirio 2012   Acanthodactylus longipes Long-footed Fringe-toed Lizard Trape, Trape & Chirio 2012 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Dunger 1967 Papenfuss 1969 Szczerbak 1975 Trape, Trape & Chirio 2012 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Arnold 1989 Dunger 1967 Reid 1986 Romer 1953 Trape, Trape & Chirio 2012 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Dunger 1967 Papenfuss 1969 Trape, Trape & Chirio 2012  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus togoensis Sahelian Fan-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 [as P. ragazzii] Butler 1986 [as P. h. hasselquistii] Dunger 1968 [as P. h. hasselquistii] Heimes 1987 [as P. ragazzii] Joger 1981 [as P. hasselquistii] Sindaco & Jeremchenko 2008 [as P. ragazzii] Trape, Trape & Chirio 2012 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola ephippiata African Wall Gecko Dunger 1968 Joger 1984 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides delislei Central Saharan Three-fingered Skink Dunger 1972   Chalcides thierryi Togo Cylindrical Skink Carranza, Arnold, Geniez, Roca & Mateo 2008 Dunger 1972 Greenbaum 2005 Papenfuss 1969 Pasteur 1981 Trape, Trape & Chirio 2012 Genus Feylinia (African Snake Skinks)   Feylinia currori Western Forest Snake Skink Broadley & Howell 1991 Brygoo & Roux-Esteve 1983 Dunger 1972 Trape, Trape & Chirio 2012 Genus Leptosiaphos (Slender Leaf-litter Skinks)   Leptosiaphos aloysiisabaudiae Uganda Five-toed Skink Broadley 1989 Schmitz, Ineich & Chirio 2005   Leptosiaphos dungeri Nigerian Leaf-litter Skink Trape, Trape & Chirio 2012 Genus Melanoseps (Black Limbless Skinks)   Melanoseps occidentalis Western Limbless Skink Brygoo & Roux-Esteve 1981 Romer 1953 Trape, Trape & Chirio 2012 Genus Mochlus (African Supple Skinks)   Mochlus fernandi Western Red-flanked Skink Trape, Trape & Chirio 2012 Wagner, Boehme, Pauwels & Schmitz 2009   Mochlus guineensis Guinean Supple Skink Butler 1986 Dunger 1973 Trape, Trape & Chirio 2012   Mochlus mocquardi Chad Supple Skink Broadley 1966 [as sundevalli] Butler 1986 [as sundevalli] Dunger 1973 [as sundevalli] Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis breviceps Short-headed Snake-eyed Skink Reid 1986 Trape, Trape & Chirio 2012   Panaspis togoensis Togo Snake-eyed Skink Boehme & Nikolaus 1989 Butler 1986 Dunger 1973 Fuhn 1972 Medina, Bauer, Branch, Schmitz, Conradie, Nagy, Hibbitts, Ernst, Portik, Nielsen, Colston, Kusamba, Behangana, Roedel & Greenbaum 2016 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Perret 1973 Reid 1986 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012 Genus Scincopus (Tiger Skinks)   Scincopus fasciatus Banded Tiger Skink Dunger 1973 Lanza & Corsi 1981 Papenfuss 1969 Trape, Trape & Chirio 2012 Genus Scincus (Sand Fish)   Scincus albifasciatus White-banded Sand Fish Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Butler 1986 Dunger 1972 Hoogmoed 1974 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Reid 1986 Romer 1953 [as Mabuya blandingi] Trape, Trape & Chirio 2012   Trachylepis maculilabris Common Speckled-lipped Skink Butler 1986 Dunger 1972 Hoogmoed 1974 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Reid 1986 Romer 1953 Trape, Trape & Chirio 2012   Trachylepis perrotetii Orange-flanked Skink Butler 1986 Cansdale 1955 Dunger 1972 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape, Trape & Chirio 2012   Trachylepis polytropis Greater Forest Skink Reid 1986 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Butler 1986 Dunger 1972 Joger 1981 Koppetsch 2020 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape, Trape & Chirio 2012   Trachylepis rodenburgi Ghanese Savanna Skink Butler 1986 Hoogmoed 1974 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Papenfuss 1969 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Boehme & Ziegler 1997 Butler 1986 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape, Trape & Chirio 2012   Varanus stellatus African Forest Monitor Bayless 2002 [as V. ornatus] Boehme & Ziegler 1997 [as V. ornatus] Boehme & Ziegler in Pianka, King & King (eds.) 2004 [as V. ornatus] Reid 1986 [as V. ornatus] Trape, Trape & Chirio 2012 [as V. ornatus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Akinpelu & Areo 2007 Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Trape & Balde 2014 Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Butler & Reid 1990 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hippocrepis West African Herald Snake Chirio & LeBreton 2007 Rasmussen, Chirio & Ineich 2000 Trape & Balde 2014   Crotaphopeltis hotamboeia Common Herald Snake Akinpelu & Areo 2007 Butler & Reid 1990 Hughes 1983 Johnsen 1962 Trape & Balde 2014 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Bates & Broadley 2018 Bates 2013 Chirio & LeBreton 2007 Trape & Balde 2014   Dasypeltis fasciata Central African Egg Eater Bates & Broadley 2018 Butler & Reid 1990 Chirio & LeBreton 2007 Gans 1959 Hughes 1983 Romer 1953 Trape & Balde 2014   Dasypeltis gansi Savanna Egg Eater Bates, Trape & Spawls 2013 Chirio & LeBreton 2007 Trape & Balde 2014 Trape & Mane 2006   Dasypeltis parascabra Guinean Egg Eater Chippaux & Jackson 2019 Trape & Balde 2014 Trape, Mediannikov & Trape 2012   Dasypeltis sahelensis Sahel Egg Eater Chippaux & Jackson 2019 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa duchesnei Blue-tailed Nocturnal Tree Snake Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Rasmussen 1989 Romer 1953 Trape & Balde 2014   Dipsadoboa guineensis Guinean Nocturnal Tree Snake Rasmussen 1989 [as brevirostris] Trape & Balde 2014   Dipsadoboa underwoodi Cameroon Nocturnal Tree Snake Chirio & LeBreton 2007   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Butler & Reid 1990 Hughes 1983 Rasmussen 1993 Trape & Balde 2014   Dipsadoboa viridis Green Nocturnal Tree Snake Jaekel 1985 [as elongata] Rasmussen 1993 Trape & Balde 2014   Dipsadoboa weileri Grey Nocturnal Tree Snake Branch, Conradie & Tolley in Branch, Bayliss, Bittencourt-Silva, Conradie, Engelbrecht, Loader, Menegon, Nanvonamuquitxo & Tolley 2019 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) Butler & Reid 1990 [as D. typus] David & Ineich 1999 [as D. typus] Hughes 1983 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Balde 2014 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014   Grayia tholloni Nandi Water Snake Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Butler & Reid 1990 Dunger 1973 Hughes 1983 Trape & Balde 2014   Hapsidophrys smaragdinus Common Emerald Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1973 Hughes 1983 Romer 1953 Trape & Balde 2014 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014   Meizodon plumbiceps Black-headed Smooth Snake Hughes 1983   Meizodon regularis East African Crowned Smooth Snake Butler & Reid 1990 Trape & Balde 2014 Genus Natriciteres (African Marsh Snakes)   Natriciteres fuliginoides Collared Marsh Snake Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014   Natriciteres olivacea Olive Marsh Snake Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014   Natriciteres variegata Variable Marsh Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1972 Hughes 1983 Trape & Balde 2014 Genus Philothamnus (African Green Snakes)   Philothamnus carinatus Barred Green Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014   Philothamnus heterodermus Emerald Green Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1973 Hughes 1983 Trape & Balde 2014   Philothamnus heterolepidotus Slender Green Snake Butler & Reid 1990 Dunger 1973 Hughes 1983 Trape & Balde 2014   Philothamnus irregularis Irregular Green Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1973 Hughes 1983 Trape & Balde 2014   Philothamnus nitidus Cameroon Green Snake Butler & Reid 1990 Dunger 1973 Hughes 1983 Trape & Balde 2014   Philothamnus semivariegatus Variegated Bush Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1973 Hughes 1983   Philothamnus smithii Smith's Bush Snake Trape & Balde 2014 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps perreti Cameroon Whip Snake Butler & Reid 1990 [as florulentus] Chippaux & Jackson 2019 Hughes 1983 [as florulentus] Schatti 1988 Sindaco, Venchi & Grieco 2013 Trape & Balde 2014 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus tripolitanus North African Cat Snake Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Schleich, Kastle & Kabisch 1996 [as obtusus] Sindaco, Venchi & Grieco 2013 Trape & Balde 2014 Wilms, Wagner & Spawls 2021   Telescopus variegatus Variable Cat Snake Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Trape & Balde 2014 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Broadley 2001 Butler & Reid 1990 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Romer 1953 Spawls & Branch 1995 Trape & Balde 2014 Genus Thrasops (Black Tree Snakes)   Thrasops flavigularis Yellow-throated Black Tree Snake Broadley & Wallach 2002 Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Trape & Balde 2014 Villiers 1975   Thrasops jacksonii Common Black Tree Snake Trape & Balde 2014   Thrasops occidentalis Western Black Tree Snake Broadley & Wallach 2002 Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Villiers 1975 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas adamanteus Eastern Powdered Cat Snake Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Akinpelu & Areo 2007 Butler & Reid 1990 Chippaux & Jackson 2019 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014   Toxicodryas pulverulenta Western Powdered Cat Snake Akinpelu & Areo 2007 Butler & Reid 1990 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Hughes 1983 Romer 1953 Trape & Balde 2014  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014   Dendroaspis viridis Western Green Mamba Akinpelu & Areo 2007 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Trape & Balde 2014 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1971 Butler & Reid 1990 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Genus Naja (Typical Cobras)   Naja annulata Ringed Water Cobra Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 Trape & Balde 2014 [as N. boulengerina (sic)]   Naja haje Egyptian Cobra Sindaco, Venchi & Grieco 2013 Trape & Balde 2014 Trape, Chirio, Broadley & Wuster 2009   Naja katiensis Brown Spitting Cobra Broadley 1968 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Naja melanoleuca Central African Forest Cobra Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja nigricollis Black-necked Spitting Cobra Akinpelu & Areo 2007 Butler & Reid 1990 Hughes 1983 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Romer 1953 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Naja savannula West African Banded Cobra Chirio & LeBreton 2007 [as N. subfulva] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Chippaux & Jackson 2019 Trape & Balde 2014 Trape, Chirio, Broadley & Wuster 2009 Genus Pseudohaje (Tree Cobras)   Pseudohaje goldii Central African Tree Cobra Butler & Reid 1990 Chippaux & Jackson 2019 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Villiers 1975   Pseudohaje nigra Black Tree Cobra Butler & Reid 1990 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Butler & Reid 1990 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus lineatus Lined Centipede Eater Akinpelu & Areo 2007   Aparallactus lunulatus Reticulated Centipede Eater Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Aparallactus modestus Western Forest Centipede Eater Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake Boulenger 1896 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis corpulenta Fat Stiletto Snake Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake Butler & Reid 1990 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake Butler & Reid 1990 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Laurent 1950 Spawls & Branch 1995 Trape & Balde 2014 Villiers 1975 Wallach, Williams & Boundy 2014   Atractaspis micropholis Sahelian Stiletto Snake Berre 1989 Butler & Reid 1990 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Papenfuss 1969 Trape & Balde 2014 Trape & Mane 2006 Trape, Mane & Ineich 2006 Villiers 1975 Wallach, Williams & Boundy 2014   Atractaspis reticulata Reticulate Stiletto Snake Butler & Reid 1990 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis watsoni Sokoto Stiletto Snake Butler & Reid 1990 [as A. microlepidota] Chirio & LeBreton 2007 David & Ineich 1999 [as A. microlepidota] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as A. m. microlepidota] Hughes 1983 [as A. microlepidota] Spawls & Branch 1995 [as A. microlepidota] Trape & Balde 2014 Trape, Mane & Ineich 2006 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Johnsen 1962 Trape & Balde 2014   Boaedon lineatus Lined House Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Romer 1953 Trape & Balde 2014   Boaedon olivaceus Olive House Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Trape & Balde 2014   Boaedon paralineatus Central African Lined House Snake Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019   Boaedon perisilvestris Central African Savanna House Snake Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020   Boaedon virgatus West African House Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Romer 1953 Trape & Balde 2014 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus lineatus Red-and-Black Striped Snake Akinpelu & Areo 2007 Butler & Reid 1990 Chirio & LeBreton 2007 Hughes 1983 Trape & Balde 2014 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Genus Dendrolycus (Central African Banded Snake)   Dendrolycus elapoides Central African Banded Snake Butler & Reid 1990 Hughes 1983 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Butler & Reid 1990 Hughes 1983 Hughes 2004 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Trape & Balde 2014 Villiers 1975   Dromophis praeornatus Ornate Olympic Snake Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Papenfuss 1969 Trape & Balde 2014 Villiers 1975 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Trape & Balde 2014   Gonionotophis klingi Western Forest Dwarf File Snake Butler & Reid 1990 Chippaux & Jackson 2019 Dunger 1971 Hughes 1983 Trape & Balde 2014 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Hughes 1983 Trape & Balde 2014 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Akinpelu & Areo 2007 Butler & Reid 1990 Chippaux & Jackson 2019 Dunger 1971 Hughes 1983 Papenfuss 1969 Romer 1953 Trape & Balde 2014 Villiers 1975   Limaformosa guirali White-speckled Forest File Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1971 Hughes 1983 Romer 1953 Trape & Balde 2014 Genus Lycophidion (African Wolf Snakes)   Lycophidion irroratum Western Forest Wolf Snake Butler & Reid 1990 Hughes & Barry 1969 Hughes 1983 Trape & Balde 2014   Lycophidion laterale Masked Wolf Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014   Lycophidion multimaculatum Many-spotted Wolf Snake Butler & Reid 1990 [as L. capense] Hughes 1983 [as L. capense] Trape & Balde 2014   Lycophidion nigromaculatum Black-spotted Wolf Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014   Lycophidion ornatum Ornate Wolf Snake Broadley 1996 Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Trape & Balde 2014   Lycophidion semicinctum Semi-annulated Wolf Snake Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Genus Mehelya (Greater File Snakes)   * Mehelya egbensis Egbe File Snake Butler & Reid 1990 Chippaux & Jackson 2019 Dunger 1971 Hughes 1983 Trape & Balde 2014   Mehelya poensis Western Forest File Snake Akinpelu & Areo 2007 Butler & Reid 1990 Dunger 1971 Hughes 1983 Romer 1953 Trape & Balde 2014   Mehelya stenophthalmus Small-eyed File Snake Butler & Reid 1990 Dunger 1971 Hughes 1983 Trape & Balde 2014 Genus Polemon (African Snake Eaters)   Polemon acanthias Five-striped Snake Eater Butler & Reid 1990 Chippaux & Jackson 2019 Doucet 1963 Hughes 1983 Trape & Balde 2014 Villiers 1975   Polemon collaris Fawn-headed Snake Eater Butler & Reid 1990 Chippaux & Jackson 2019 Pitman 1974 Trape & Balde 2014 Villiers 1975   Polemon gabonensis Gabon Snake Eater Butler & Reid 1990 Chippaux & Jackson 2019 Hughes 1983 Trape & Balde 2014   Polemon gracilis Slender Snake Eater Chippaux & Jackson 2019 Chirio & LeBreton 2007 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chirio, Ineich, Schmitz & Trape 2011   Prosymna greigerti Sudan Savanna Shovel-snout Butler & Reid 1990 Chirio & LeBreton 2007 Chirio, Ineich, Schmitz & Trape 2011 Hughes 1983 [as P. meleagris] Papenfuss 1969 [as P. laurenti]   Prosymna meleagris Speckled Shovel-snout Broadley 1980 Butler & Reid 1990 Chirio, Ineich, Schmitz & Trape 2011 Hughes 1983 Trape & Balde 2014 Trape & Mane 2006 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Butler & Reid 1990 [as P. sibilans] Hughes 1983 [as P. sibilans] Romer 1953 [as P. sibilans] Trape & Balde 2014 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Akinpelu & Areo 2007 Butler & Reid 1990 Hughes 1983 Joger 1981 Papenfuss 1969 Trape & Balde 2014   Psammophis mossambicus East African Olive Grass Snake Chirio & LeBreton 2007 [as occidentalis] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis phillipsii West African Olive Grass Snake Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 Butler & Reid 1990 Hughes 1983 Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis sudanensis Sudan Sand Snake Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Butler & Reid 1990 Chirio & Ineich 1991 Loveridge 1957 Pitman 1974 Trape & Balde 2014  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis macrorhyncha Large-snouted Thread Snake Butler & Reid 1990 Gasperetti 1988 Hahn 1978 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 [as "cf. macrochynchus"]   Myriopholis narirostris Sahel Thread Snake Butler & Reid 1990 Hahn 1980 Hughes 1983 McDiarmid, Campbell & Toure 1999 Papenfuss 1969 Trape & Balde 2014 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma bicolor Bicoloured Thread Snake Akinpelu & Areo 2007 Butler & Reid 1990 Hahn & Wallach 1998 Hahn 1980 [as Leptotyphlops brevicaudus] Hughes 1983 McDiarmid, Campbell & Toure 1999 [as brevicaudus] Papenfuss 1969 [as brevicauda] Trape & Balde 2014   * Tricheilostoma greenwelli Ibadan Thread Snake Chippaux & Jackson 2019 Trape & Balde 2014 Wallach & Boundy 2005   Tricheilostoma sundewalli Central African Thread Snake Hughes 1983 Trape & Balde 2014  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Butler & Reid 1990 Hughes 1983 McDiarmid, Campbell & Toure 1999 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Papenfuss 1969 Romer 1953 Trape & Balde 2014   Python sebae Northern Rock Python Akinpelu & Areo 2007 Broadley 1984 Butler & Reid 1990 Hughes 1983 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Spawls & Branch 1995 Trape & Balde 2014 Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops congestus Marbled Blind Snake Broadley & Wallach 2009 Chippaux & Jackson 2019 Hahn 1980 McDiarmid, Campbell & Toure 1999 Spawls, Howell, Drewes & Ashe 2002 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops lineolatus Lineolate Blind Snake Butler & Reid 1990 Hahn 1980 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Akinpelu & Areo 2007 Butler & Reid 1990 Hahn 1980 Hughes 1983 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops steinhausi Common Cameroon Blind Snake Broadley & Wallach 2009 Butler & Reid 1990 Chippaux & Jackson 2019 Hahn 1980 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Letheobia (African Gracile Blind Snakes)   Letheobia crossii Nigerian Gracile Blind Snake Butler & Reid 1990 Chippaux & Jackson 2019 Hahn 1980 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014   Letheobia weidholzi Poli Gracile Blind Snake Chippaux & Jackson 2019 [as praeocularis] Rasmussen 1997 [as praeocularis] Trape & Balde 2014 [as praeocularis] Trape, Kodindo, Djiddi, Mad-Toingue & Kerah 2020  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   Atheris broadleyi Lipondji Bush Viper Chippaux & Jackson 2019   Atheris chlorechis West African Green Bush Viper Butler & Reid 1990 David & Ineich 1999 Hughes 1983 McDiarmid, Campbell & Toure 1999 Romer 1953 Spawls & Branch 1995 Trape & Balde 2014   Atheris squamigera Southern Green Bush Viper Butler & Reid 1990 Chippaux & Jackson 2019 David & Ineich 1999 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Butler & Reid 1990 David & Ineich 1999 Hughes 1983 McDiarmid, Campbell & Toure 1999 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Spawls & Branch 1995 Trape & Balde 2014   Bitis gabonica Gaboon Adder Akinpelu & Areo 2007 Broadley 1983 Butler & Reid 1990 Hughes 1983 Romer 1953 Spawls & Branch 1995 Trape & Balde 2014   Bitis nasicornis African Nose-horned Viper Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014   Bitis rhinoceros Rhinoceros Viper Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Genus Causus (Night Adders)   Causus lichtensteinii Forest Night Adder Akani, Luiselli, Tooze, Angelici, Corti & Zuffi 2001 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Spawls & Branch 1995 Trape & Balde 2014   Causus maculatus West African Night Adder Akani, Luiselli, Tooze, Angelici, Corti & Zuffi 2001 Akinpelu & Areo 2007 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Joger 1981 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Spawls & Branch 1995 Trape & Balde 2014   Causus resimus Green Night Adder Akani, Luiselli, Tooze, Angelici, Corti & Zuffi 2001 David & Ineich 1999 Hughes 1983 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014   Causus rhombeatus Rhombic Night Adder Akani, Luiselli, Tooze, Angelici, Corti & Zuffi 2001 Akinpelu & Areo 2007 Butler & Reid 1990 David & Ineich 1999 Hughes 1983 Johnsen 1962 Romer 1953 Spawls & Branch 1995 Trape & Balde 2014 Genus Echis (Carpet Vipers)   Echis leucogaster White-bellied Carpet Viper Butler & Reid 1990 David & Ineich 1999 Hughes 1976 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Trape & Balde 2014   Echis ocellatus West African Carpet Viper Butler & Reid 1990 David & Ineich 1999 Hughes 1976 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Spawls & Branch 1995 Trape & Balde 2014   Echis romani Chadian Carpet Viper Trape 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle King & Burke 1989 Mettler, Clyde-Brockway, Honarvar & Paladino 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boycott & Bourquin 2008 [as subrufa] Fritz & Havas 2006 [as subrufa] Iverson 1992 [as subrufa] Joger 1981 [as subrufa] King & Burke 1989 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelomedusa subrufa Common Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelomedusa variabilis West African Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Bour 2008 King & Burke 1989 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 King & Burke 1989 McCord, Joseph-Ouni & Bour 2004 Nneji, Adeola, Okeyoyin, Oladipo, Saidu, Samuel, Usongo, Adedeji, Omotoso, Adeyi, Ugwumba & Ugwumba 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Pelusios gabonensis Forest Mud Turtle Chirio & LeBreton 2007   Pelusios niger West African Black Mud Turtle Chirio & LeBreton 2007 King & Burke 1989 Luiselli, Bour, Petrozzi, Akani & Segniagbeto 2018 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise King & Burke 1989 Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Kinixys (Hinge-back Tortoises)   Kinixys erosa Forest Hinge-back Tortoise Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 King & Burke 1989 Luiselli & Diagne 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys homeana West African Hinge-back Tortoise Akani, Luiselli, Ogbeibu, Onwuteaka, Chuku, Osakwe, Bombi, Amuzie, Uwagbae & Guo 2010 King & Burke 1989 Luiselli & Diagne 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys nogueyi Western Savanna Hinge-back Tortoise Chirio & LeBreton 2007 King & Burke 1989 [as belliana] McCord, Joseph-Ouni & Tabaka 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 King & Burke 1989 McCord & Joseph-Ouni 2003 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977   Cyclanorbis senegalensis Senegal Flapshell Turtle King & Burke 1989 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Papenfuss 1969 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012