Reptiles of
Tocantins [Brazil]

Related checklists:  Brazil

Bibliography of the reptiles of Tocantins Taxonomic history of the reptiles of Tocantins Introduced species
Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates
Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Worm Lizards (Amphisbaenia)
= endemicEX = extinct or assumed extinct

Species
References
 Family Amphisbaenidae (Typical Worm Lizards)
Genus Amphisbaena (Neotropical Worm Lizards)
 
Amphisbaena alba

Common Red Worm Lizard

Costa, Guedes & Bernils 2021 Mott & Vieites 2009 Mott 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019
 
Amphisbaena anaemariae

Serra da Mesa Worm Lizard

Costa, Guedes & Bernils 2021
 
Amphisbaena filiformis

Babaculandia Worm Lizard

Costa, Guedes & Bernils 2021 Ribeiro, Gomes, Silva, Cintra & Silva 2016
 
Amphisbaena leeseri

Urucum Worm Lizard

Costa, Guedes & Bernils 2021 Mott & Vieites 2009 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019
 
Amphisbaena miringoera

Rio Tapirapes Worm Lizard

Costa, Guedes & Bernils 2021
 
Amphisbaena silvestrii

Southern Mato Grosso Worm Lizard

Costa, Guedes & Bernils 2021
 
Amphisbaena talisiae

Serra da Pitomba Worm Lizard

Costa, Guedes & Bernils 2021
 
Amphisbaena vermicularis

Vermiculate Worm Lizard

Costa, Guedes & Bernils 2021 Gans & Amdur 1966 Mott & Vieites 2009 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro, Gomides & Costa 2020 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019
Genus Anops (Keel-headed Worm Lizards)
  *
Anops acrobeles

Jalapao Worm Lizard

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro, Castro-Mello & Nogueira 2009
Genus Aulura (Auro Worm Lizard)
 
Aulura anomala

Auro Worm Lizard

Costa, Guedes & Bernils 2021
Genus Bronia (Brazilian Worm Lizards)
  *
Bronia kraoh

Godias Worm Lizard

Castro-Mello 2003 Costa, Guedes & Bernils 2021 Gans 2005 Mott & Vieites 2009 Pinna, Mendonca, Bocchiglieri & Fernandes 2010 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019 Vanzolini 1971
  *
Bronia saxosa

Tocantins Worm Lizard

Castro-Mello 2003 Costa, Guedes & Bernils 2021 Mott & Vieites 2009 Pinna, Mendonca, Bocchiglieri & Fernandes 2010 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019
Genus Cercolophia (Keel-tailed Worm Lizards)
 
Cercolophia roberti

Ipiranga Worm Lizard

Costa, Guedes & Bernils 2021 Mott & Vieites 2009 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019
Genus Leposternon (South American Worm Lizards)
 
Leposternon infraorbitale

Central Brazilian Worm Lizard

Costa, Guedes & Bernils 2021
 
Leposternon polystegum

East Brazilian Worm Lizard

Costa, Guedes & Bernils 2021 Mott & Vieites 2009 Mott 2006 Ribeiro, Nogueira, Cintra, Silva & Zaher 2011 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019 Ribeiro, Vaz-Silva & Santos 2008 Tavares & Ribeiro 2014

Crocodilians (Crocodylia)
= endemicEX = extinct or assumed extinct

Species
References
 Family Alligatoridae (Alligators and Caimans)
Genus Caiman (Typical Caimans)
 
Caiman crocodilus

Common Caiman

Costa, Guedes & Bernils 2021 Dornas, Silva, Silva & Malvasio 2021
Genus Melanosuchus (Black Caiman)
 
Melanosuchus niger

Black Caiman

Costa Pereira & Malvasio 2014 Costa, Guedes & Bernils 2021 Dornas, Silva, Silva & Malvasio 2021
Genus Paleosuchus (Dwarf Caimans)
 
Paleosuchus palpebrosus

Cayenne Dwarf Caiman

Costa, Guedes & Bernils 2021 Dornas, Silva, Silva & Malvasio 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Paleosuchus trigonatus

Northwestern Dwarf Caiman

Costa, Guedes & Bernils 2021

Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Anolidae (Anoles)
Genus Norops (Brilliant Anoles)
 
Norops brasiliensis

Brazilian Golden-scaled Anole

Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as chrysolepis] Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015 Vitt, Shepard, Vieira, Caldwell, Colli & Mesquita 2008
 
Norops meridionalis

Paraguayan Anole

Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 
Norops ortonii

Amazon Bark Anole

Costa, Guedes & Bernils 2021 d'Assuncao, Filho, Santos Araujo & Moura 2012
 Family Gekkonidae (Typical Geckos)
Genus Hemidactylus (Half-toed Geckos)
 
Hemidactylus brasilianus

Brazilian Half-toed Gecko

Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 Family Gymnophthalmidae (Scaly-tongued Spectacled Tejus)
Genus Bachia (Earless Tejus)
 
Bachia bresslaui

Sao Paulo Earless Teju

Costa, Guedes & Bernils 2021
  *
Bachia micromela

Guarai Earless Teju

Costa, Guedes & Bernils 2021 Rodrigues, Pavan & Curcio 2007 Teixeira, Recoder, Camacho, Sena, Navas & Rodrigues 2013
  *
Bachia oxyrhina

Mateiros Earless Teju

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Rodrigues, Camacho, Nunes, Recoder, Teixeira, Valdujo, Ghellere, Mott & Nogueira 2008 Teixeira, Recoder, Camacho, Sena, Navas & Rodrigues 2013
  *
Bachia psamophila

Porto Nacional Earless Teju

Costa, Guedes & Bernils 2021 Rodrigues, Pavan & Curcio 2007 Teixeira, Recoder, Camacho, Sena, Navas & Rodrigues 2013
Genus Cercosaura (Eastern Shade Tejus)
 
Cercosaura olivacea

East Brazilian Shade Teju

Costa, Guedes & Bernils 2021 Dantas Sales, Silva Jorge, Meira-Ribeiro & Xavier Freire 2014 [as ocellata] Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as ocellata] Ribeiro-Junior & Amaral 2016 [as "C. ocellata ssp."] Sturaro, Rodrigues, Colli, Knowles & Avila-Pires 2018
 
Cercosaura schreibersii

South Brazilian Shade Teju

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
Genus Colobosaura (Short-thumbed Spectacled Tejus)
 
Colobosaura modesta

Morro da Garcza Spectacled Teju

Costa, Guedes & Bernils 2021 Freire, Jorge & Ribeiro 2012 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
Genus Loxopholis (Western Root Tejus)
 
Loxopholis percarinata

Striped Root Teju

Brunes, Silva, Marques-Souza, Rodrigues & Pellegrino 2019 Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
Genus Micrablepharus (Snake-eyed Glossy Tejus)
 
Micrablepharus atticolus

Cerrado Snake-eyed Glossy Teju

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
 
Micrablepharus maximiliani

Blue-tailed Snake-eyed Glossy Teju

Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Moura, Dayrell & Avelar Sao Pedro 2010 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
Genus Neusticurus (Guianan Water Tejus)
 
Neusticurus bicarinatus

Two-keeled Water Teju

Costa, Guedes & Bernils 2021 Marques-Souza, Prates, Fouquet, Camacho, Kok, Nunes, Vechio, Recoder, Mejia, Teixeira-Junior, Barrio-Amoros, Cassimiro, Lima, Sena & Rodrigues 2018 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior & Amaral 2017
Genus Vanzosaura (Red-tailed Spectacled Tejus)
 
Vanzosaura savanicola

Cerrado Red-tailed Spectacled Teju

Colli, Giugliano, Mesquita & Franca 2009 [as rubricauda] Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 [as rubricauda] Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as rubricauda] Recoder, Ribeiro & Rodrigues 2013 [as rubricauda] Recoder, Werneck, Teixeira, Colli, Sites & Rodrigues 2014
 Family Hoplocercidae (Club-tailed Iguanas)
Genus Hoplocercus (Eastern Club-tailed Iguana)
 
Hoplocercus spinosus

Eastern Club-tailed Iguana

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 Family Iguanidae (Greater Iguanas)
Genus Iguana (Green Iguanas)
 
Iguana iguana

South American Green Iguana

Avila-Pires 1995 Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 Family Phyllodactylidae (Leaf-toed Geckos)
Genus Gymnodactylus (Naked-toed Geckos)
 
Gymnodactylus amarali

Maranhao Naked-toed Gecko

Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Gamble, Colli, Rodrigues, Werneck & Simons 2012 Mesquita, Colli, Franca & Vitt 2006 [as carvalhoi] Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015 Vanzolini 2004 Vanzolini 2005 [as carvalhoi]
Genus Phyllopezus (South American Robust Geckos)
 
Phyllopezus pollicaris

Common Robust Gecko

Condez, Tonini, Pereira-Ribeiro & Dubeux 2021 Costa, Guedes & Bernils 2021 Gamble, Colli, Rodrigues, Werneck & Simons 2012 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 Family Polychrotidae (Monkey Lizards)
Genus Polychrus (Monkey Lizards)
 
Polychrus acutirostris

Sharp-nosed Monkey Lizard

Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 
Polychrus marmoratus

Marbled Monkey Lizard

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 Family Scincidae (Skinks)
Genus Aspronema (South American White-striped Skinks)
 
Aspronema dorsivittatum

South American White-striped Skink

Costa, Guedes & Bernils 2021
Genus Brasiliscincus (Brazilian Small-headed Skinks)
 
Brasiliscincus heathi

Caatinga Small-headed Skink

Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. heathi"]
Genus Copeoglossum (Neotropical Spotted Skinks)
 
Copeoglossum nigropunctatum

Black-spotted Skink

Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Miralles, Barrio-Amoros, Rivas & Chaparro-Auza 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. nigropunctatum"] Ribeiro-Junior & Amaral 2016
Genus Manciola (South American Small-handed Skink)
 
Manciola guaporicola

South American Small-handed Skink

Costa, Guedes & Bernils 2021 Hedges & Conn 2012 Miralles & Carranza 2010 Ribeiro-Junior & Amaral 2016
Genus Notomabuya (Southern Neotropical Skink)
 
Notomabuya frenata

Southern Neotropical Skink

Caldeira Costa, Dias Fernandes, Vrcibradic & Neves Feio 2008 Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Hedges & Conn 2012 Miralles & Carranza 2010 Pinto & Araujo 2000 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Vrcibradic, Almeida-Gomes, Borges-Junior, Kiefer, Sluys & Raocha 2006
 Family Sphaerodactylidae (Least Geckos, etc.)
Genus Coleodactylus (Sheath-toed Pygmy Geckos)
 
Coleodactylus brachystoma

Cerrado Pygmy Gecko

Costa, Guedes & Bernils 2021 Freitas, Franca & Oliveira-Lima 2011 Geurgas, Rodrigues & Moritz 2008 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Coleodactylus meridionalis

Atlantic Forest Pygmy Gecko

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
Genus Gonatodes (American Padless Geckos)
 
Gonatodes humeralis

Bridled Forest Gecko

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 Family Teiidae (Greater Tejus)
Genus Ameiva (South American Ameivas)
 
Ameiva ameiva

South American Giant Ameiva

Avila-Pires 1995 Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Ribeiro-Junior & Amaral 2016 Vanzolini 1972
Genus Ameivula (Eastern South American Whiptails)
 
Ameivula jalapensis

Jalapao Whiptail

Arias, Carvalho, Rodrigues & Zaher 2011 Arias, Carvalho, Zaher & Rodrigues 2014 Arias, Recoder, Alvarez, Etchepare, Quipildor, Lobo & Rodrigues 2018 Arias, Teixeira Junior, Recoder, Morato de Carvalho, Zaher & Rodrigues 2014 Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Harvey, Ugueto & Gutberlet 2012 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Ameivula mumbuca

Mateiros Whiptail

Arias, Carvalho, Rodrigues & Zaher 2011 Arias, Carvalho, Zaher & Rodrigues 2014 Arias, Recoder, Alvarez, Etchepare, Quipildor, Lobo & Rodrigues 2018 Arias, Teixeira Junior, Recoder, Morato de Carvalho, Zaher & Rodrigues 2014 Colli, Caldwell, Costa, Gainsbury, Garda, Mesquita, Filho, Soares, Silva, Valdujo, Vieira, Vitt, Werneck, Wiederhecker & Zatz 2003 Costa, Guedes & Bernils 2021 Harvey, Ugueto & Gutberlet 2012 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Ameivula ocellifera

Brazilian Ocellated Whiptail

Aparicio, Avila-Pires, Moravec & Nogueira 2019 Costa, Guedes & Bernils 2021 Mesquita & Colli 2003
Genus Kentropyx (Spurred Whiptails)
 
Kentropyx calcarata

Striped Forest Whiptail

Costa, Guedes & Bernils 2021 Gallagher & Dixon 1992 Ribeiro-Junior & Amaral 2016
 
Kentropyx paulensis

Sao Paulo Whiptail

Costa, Guedes & Bernils 2021 Gallagher & Dixon 1992 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. paulensis"]
Genus Salvator (Southern Tejus)
 
Salvator duseni

Parana Teju

Campos, Oda, Custodio, Koppe & Dartora 2011 Costa, Guedes & Bernils 2021
 
Salvator merianae

Southern Black-and-White Teju

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016
Genus Tupinambis (Northern Tejus)
 
Tupinambis quadrilineatus

Four-striped Teju

Abrahao-Morato, Meer, Bornschein, Vieira Capela, Almeida Ulandowski & Zampier 2015 Costa, Guedes & Bernils 2021 Freitas, Oliveira-Lima & Franca 2011 Koehler & Langerwerf 2000 Manzani & Abe 1997 Mesquita, Colli, Franca & Vitt 2006 Ribeiro-Junior & Amaral 2016 Silva, Ribeiro-Junior & Avila-Pires 2018 Silveira 2009
 
Tupinambis teguixin

Amazonian Black-and-White Teju

Avila-Pires 1995 Colli, Peres & Cunha 1998 Costa, Guedes & Bernils 2021 Harvey, Ugueto & Gutberlet 2012 Murphy, Jowers, Lehtinen, Charles, Colli, Peres, Hendry & Pyron 2016 [based on map] Passos, Lima-Araujo, Melo & Borges-Nojosa 2013 Ribeiro-Junior & Amaral 2016 Silva, Ribeiro-Junior & Avila-Pires 2018
 Family Tropiduridae (Rough-tailed Iguanas)
Genus Plica (Tree Runners)
 
Plica umbra

Smooth-necked Tree Runner

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
Genus Stenocercus (Whorl-tailed Iguanas)
 
Stenocercus dumerilii

Para Whorl-tailed Iguana

Costa, Guedes & Bernils 2021
 
Stenocercus quinarius

Five-crested Whorl-tailed Iguana

Costa, Guedes & Bernils 2021
Genus Tropidurus (Eastern Keel-tailed Iguanas)
 
Tropidurus hispidus

Spotted Keel-tailed Iguana

Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015
 
Tropidurus insulanus

Cachimbo Keel-tailed Iguana

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015
 
Tropidurus itambere

Sorocaba Keel-tailed Iguana

Carvalho 2013 Costa, Guedes & Bernils 2021
 
Tropidurus oreadicus

Cerrado Keel-tailed Iguana

Carvalho 2013 Colli, Giugliano, Mesquita & Franca 2009 Costa, Guedes & Bernils 2021 Mesquita, Colli, Franca & Vitt 2006 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. oreadicus"] Ribeiro-Junior 2015 Rodrigues 1987
 
Tropidurus torquatus

Collared Keel-tailed Iguana

Carvalho 2013 Costa, Guedes & Bernils 2021 Kunz & Borges-Martins 2013
Genus Uranoscodon (Mop-headed Iguana)
 
Uranoscodon superciliosus

Mop-headed Iguana

Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Ribeiro-Junior 2015

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Aniliidae (South American Pipe Snake)
Genus Anilius (South American Pipe Snake)
 
Anilius scytale

South American Pipe Snake

Costa, Guedes & Bernils 2021 Silva, Souza & Puorto 2001
 Family Anomalepididae (Neotropical Blind Snakes)
Genus Liotyphlops (Smooth Blind Snakes)
 
Liotyphlops ternetzii

Paraguayan Blind Snake

Costa, Guedes & Bernils 2021 Cruz-Centeno, Swaya & Germano 2010
 Family Boidae (Typical Boas)
Genus Boa (Red-tailed Boas)
 
Boa constrictor

South American Red-tailed Boa

Costa, Guedes & Bernils 2021
Genus Corallus (Tree Boas)
 
Corallus batesii

Southern Emerald Tree Boa

Costa, Guedes & Bernils 2021
 
Corallus hortulana

Amazon Tree Boa

Costa, Guedes & Bernils 2021
Genus Epicrates (Rainbow Boas)
 
Epicrates cenchria

Common Rainbow Boa

Andrade-Junior, Soares Franca, Sudre & Passos 2020 Costa, Guedes & Bernils 2021 Passos & Fernandes 2008
 
Epicrates crassus

Paraguayan Rainbow Boa

Andrade-Junior, Soares Franca, Sudre & Passos 2020 Costa, Guedes & Bernils 2021 Passos & Fernandes 2008 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Eunectes (Anacondas)
 
Eunectes murinus

Green Anaconda

Costa, Guedes & Bernils 2021
 Family Colubridae (Typical Snakes or Colubrids)
Genus Adelphostigma (Twin-spotted Leaf-litter Snakes)
 
Adelphostigma occipitalis

Bahia Leaf-litter Snake

Costa, Guedes & Bernils 2021
Genus Apostolepis (Black-tailed Blackheads)
  *
Apostolepis adhara

Sao Salvador Blackhead

Costa, Guedes & Bernils 2021 Franca, Barbo, Silva-Junior, Silva & Zaher
 
Apostolepis flavotorquata

Yellow-collared Blackhead

Costa, Guedes & Bernils 2021 Lema & Renner 2005 Lema 2001 Lema 2002 Lima Silveira 2014 Wallach, Williams & Boundy 2014
 
Apostolepis longicaudata

Piaui Blackhead

Costa, Guedes & Bernils 2021 Curcio, Nunes, Harvey & Rodrigues 2011 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. longicaudata"] Wallach, Williams & Boundy 2014
 
Apostolepis nelsonjorgei

Minacu Blackhead

Costa, Guedes & Bernils 2021 Curcio, Nunes, Harvey & Rodrigues 2011 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Lema & Renner 2004 Lema & Renner 2012 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Wallach, Williams & Boundy 2014
 
Apostolepis polylepis

Hawkbeak Blackhead

Costa, Guedes & Bernils 2021
 
Apostolepis sanctaeritae

Tocantins Blackhead

Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Ferrarezzi, Barbo & Albuquerque 2005 [as ammodites] Lema & Renner 2007 [as ammodites] Lema & Renner 2012 [as ammodites] Nogueira, Barbo & Ferrarezzi 2012 [as ammodites] Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as ammodites] Wallach, Williams & Boundy 2014 [as ammodites]
Genus Atractus (Spindle Snakes)
 
Atractus albuquerquei

Guama River Spindle Snake

Costa, Guedes & Bernils 2021 Passos, Azevedo, Nogueira, Fernandes & Sawaya 2019
 
Atractus pantostictus

Sao Paulo Spindle Snake

Costa, Guedes & Bernils 2021 Fernandes & Puorto 1993 Passos, Fernandes, Bernils & Moura-Leite 2010 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Boiruna (Southern Snake Eaters)
 
Boiruna maculata

South American Spotted Snake Eater

Costa, Guedes & Bernils 2021
 
Boiruna sertaneja

Sertao Snake Eater

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 [as "cf. sertaneja"] Souto de Brito & Goncalves 2012
Genus Chironius (Sipos)
 
Chironius carinatus

Yellow-lipped Sipo

Costa, Guedes & Bernils 2021
 
Chironius exoletus

Green Sipo

Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993
 
Chironius flavolineatus

Yellow-striped Sipo

Albuquerque & Freitas 2011 Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993 Hamdan & Fernandes 2015 Hamdan, Scali & Fernandes 2014 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010
 
Chironius quadricarinatus

Paraguayan Sipo

Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010
 
Chironius scurrulus

Rusty Sipo

Costa, Guedes & Bernils 2021
Genus Chlorosoma (Green Bush Racers)
 
Chlorosoma viridissimum

Emerald Bush Racer

Costa, Guedes & Bernils 2021
Genus Clelia (Northern Snake Eaters)
 
Clelia clelia

Common Snake Eater

Costa, Guedes & Bernils 2021
 
Clelia plumbea

Plumbeous Snake Eater

Costa, Guedes & Bernils 2021
Genus Dipsas (Neotropical Snail Eaters)
 
Dipsas catesbyi

Ornate Snail Eater

Costa, Guedes & Bernils 2021 Lima & Prudente 2009 Sanchez-Martinez, Franca & Trevine 2022
 
Dipsas mikanii

Brazilian White-collared Snail Eater

Costa, Guedes & Bernils 2021 Giraudo & Scrocchi 2002
Genus Drymarchon (Indigo Snakes and Cribos)
 
Drymarchon corais

Yellow-tailed Cribo

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Drymoluber (Glossy Racers)
 
Drymoluber brazili

Southern Glossy Racer

Caldeira-Costa, Ribeiro-Moura & Neves-Feio 2013 Costa, Guedes & Bernils 2021
Genus Dryophylax (Northern Mock Vipers)
 
Dryophylax hypoconia

Southern Mock Viper

Costa, Guedes & Bernils 2021 Franco, Trevine, Montingelli & Zaher 2017 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021
 
Dryophylax phoenix

Petrolina Mock Viper

Costa, Guedes & Bernils 2021 Franco, Trevine, Montingelli & Zaher 2017 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021
Genus Erythrolamprus (Neotropical Smooth Snakes)
 
Erythrolamprus aesculapii

Southern Mock Coral Snake

Costa, Guedes & Bernils 2021 Curcio, Scali & Rodrigues 2015
 
Erythrolamprus almadensis

Almada Smooth Snake

Costa, Guedes & Bernils 2021
 
Erythrolamprus maryellenae

Goias Smooth Snake

Costa, Guedes & Bernils 2021 Franca 2004
 
Erythrolamprus poecilogyrus

Yellow-bellied Smooth Snake

Costa, Guedes & Bernils 2021
 
Erythrolamprus reginae

Northern Royal Smooth Snake

Ascenso, Costa & Prudente 2019 Costa, Guedes & Bernils 2021 Torres-Carvajal & Hinojosa 2020
 
Erythrolamprus taeniogaster

Banded Smooth Snake

Costa, Guedes & Bernils 2021 Dixon 1989
 
Erythrolamprus viridis

Brazilian Green Smooth Snake

Costa, Guedes & Bernils 2021 Walterman, Moraes-da-Silva & Curcio 2021
Genus Helicops (South American Keelbacks)
 
Helicops angulatus

Brown-banded Keelback

Costa, Guedes & Bernils 2021 Roberto, Ribeiro, Sousa-Delfino & Oliveira-Almeida 2009
 
Helicops leopardinus

Leopard Keelback

Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Helicops phantasma

Phantom Keelback

Costa, Guedes & Bernils 2021 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021
 
Helicops polylepis

South American Many-scaled Keelback

Costa, Guedes & Bernils 2021
 
Helicops trivittatus

Three-striped Keelback

Costa, Guedes & Bernils 2021 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021 Moraes-da-Silva, Amaro, Sales Nunes, Strussmann, Teixeira, Andrade, Sudre, Recoder, Rodrigues & Curcio 2019 Oliveira, Silva, Santos, Barriga, Gabriel & Costa-Campos 2015
Genus Hydrodynastes (False Water Cobras)
 
Hydrodynastes bicinctus

Two-banded False Water Cobra

Costa, Guedes & Bernils 2021 Franco, Fernandes & Bentim 2007
 
Hydrodynastes gigas

Giant False Water Cobra

Costa, Guedes & Bernils 2021
 
Hydrodynastes melanogigas

Black False Water Cobra

Franco, Fernandes & Bentim 2007 Santos-Junior, Adams, Buhler, Ribeiro & Carvalho 2017 Silva, Hamdan, Tonial, Silva & Cintra 2012
Genus Hydrops (South American Mud Snakes)
 
Hydrops triangularis

Red-sided Mud Snake

Albuquerque & Lema 2008 Costa, Guedes & Bernils 2021
Genus Imantodes (American Blunt-headed Tree Snakes)
 
Imantodes cenchoa

Common Blunt-headed Tree Snake

Costa, Guedes & Bernils 2021
Genus Leptodeira (Typical Cat-eyed Snakes)
 
Leptodeira annulata

South American Banded Cat-eyed Snake

Barrio-Amoros 2019 Costa, Guedes & Bernils 2021
Genus Leptophis (Neotropical Parrot Snakes)
 
Leptophis ahaetulla

Common Green Parrot Snake

Albuquerque & Fernandes 2022 Albuquerque, Santos, Borges-Nojosa & Avila 2022 Costa, Guedes & Bernils 2021 Torres-Carvajal & Teran 2021
 
Leptophis marginatus

Paraguayan Parrot Snake

Murphy, Charles, Lehtinen & Koeller 2013
Genus Lygophis (South American Striped Ground Snakes)
 
Lygophis meridionalis

Paraguayan Striped Ground Snake

Castro & Oliveira 2017 Costa, Guedes & Bernils 2021
 
Lygophis paucidens

Cerrado Striped Ground Snake

Costa, Guedes & Bernils 2021 Serrano, Vieira-Alencar, Diaz-Ricaurte & Nogueira 2020
Genus Mastigodryas (Neotropical Lizard Eaters)
 
Mastigodryas boddaerti

Brown Lizard Eater

Costa, Guedes & Bernils 2021 Montingelli 2009 Montingelli, Grazziotin, Battilana, Murphy, Zhang & Zaher 2019 Montingelli, Valencia, Benavides & Zaher 2011 Siqueira, Nascimento, Montingelli & Santos-Costa 2013
Genus Oxybelis (Neotropical Vine Snakes)
 
Oxybelis aeneus

Brown Vine Snake

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Oxybelis fulgidus

Green Vine Snake

Costa, Guedes & Bernils 2021 Scartozzoni, Salomao & Almeida-Santos 2009
Genus Oxyrhopus (Calico Snakes)
 
Oxyrhopus guibei

Parana Calico Snake

Costa, Guedes & Bernils 2021
 
Oxyrhopus melanogenys

Black-headed Calico Snake

Costa, Guedes & Bernils 2021
 
Oxyrhopus rhombifer

Rhombic Calico Snake

Costa, Guedes & Bernils 2021
 
Oxyrhopus trigeminus

Northern Brazilian Calico Snake

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Palusophis (Eastern Lizard Eater)
 
Palusophis bifossatus

Eastern Lizard Eater

Costa, Guedes & Bernils 2021 Marques & Muriel 2007 Montingelli 2009 Montingelli, Grazziotin, Battilana, Murphy, Zhang & Zaher 2019 Peters & Orejas-Miranda 1970
Genus Phalotris (South American Collared Snakes)
 
Phalotris labiomaculatus

Lip-spotted Collared Snake

Costa, Guedes & Bernils 2021 Franca, Mesquita & Garda 2005 Hamdan, Silva, Silva, Cintra & Lema 2013
Genus Philodryas (South American Bush Racers)
 
Philodryas nattereri

Paraguay Bush Racer

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Smith, Scott, Cacciali, Atkinson & Pheasey 2013 Thomas 1976
 
Philodryas olfersii

Common Green Bush Racer

Costa, Guedes & Bernils 2021
Genus Phimophis (South American Shovel-nosed Snakes)
 
Phimophis guerini

Argentine Shovel-nosed Snake

Costa, Guedes & Bernils 2021 Filho, Santana, Vieira, Alves, Montenegro & Freitas 2012 Marques, Tinoco, Browne-Ribeiro & Fazolato 2012
Genus Pseudablabes (Reticulated Bush Racers)
 
Pseudablabes patagoniensis

Patagonian Bush Racer

Costa, Guedes & Bernils 2021 Gouveia, Novelli, Vieira & Sousa 2017
Genus Pseudoboa (Neotropical Scarlet Snakes)
 
Pseudoboa coronata

Black-headed Scarlet Snake

Caldeira Costa, Cotta & MacCulloch 2015 Costa, Guedes & Bernils 2021 Zaher, Oliveira & Franco 2008
 
Pseudoboa nigra

Blackish Scarlet Snake

Costa, Guedes & Bernils 2021 Orofino, Pizzatto & Marques 2010 Zaher, Oliveira & Franco 2008
Genus Psomophis (Smooth Ground Snakes)
 
Psomophis joberti

Brazilian Ground Snake

Costa, Guedes & Bernils 2021 Myers & Cadle 1994
Genus Rodriguesophis (Collared Sharp-nosed Snakes)
 
Rodriguesophis iglesiasi

Piaui Sharp-nosed Snake

Costa, Guedes & Bernils 2021 Lima Silveira 2014 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Siphlophis (Liana Snakes)
 
Siphlophis compressus

Red-headed Liana Snake

Costa, Guedes & Bernils 2021
 
Siphlophis worontzowi

Amazon Liana Snake

Costa, Guedes & Bernils 2021 Prudente, Silva, Santos Meireles & Puorto 2017 Vechio, Teixeira-Junior, Neto & Rodrigues 2015
Genus Spilotes (Chicken Snakes)
 
Spilotes pullatus

Chicken Snake

Costa, Guedes & Bernils 2021
 
Spilotes sulphureus

Yellow-bellied Puffing Snake

Costa, Guedes & Bernils 2021
Genus Tantilla (American Centipede Snakes)
 
Tantilla melanocephala

South American Black-headed Centipede Snake

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Wilson & Mena 1980
Genus Thamnodynastes (South American Mock Vipers)
 
Thamnodynastes pallidus

Common South American Mock Viper

Costa, Guedes & Bernils 2021 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021
Genus Xenodon (Northern Mock Lanceheads and South American Hognose Snakes)
 
Xenodon merremii

Toad-eating Mock Lancehead

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Xenodon nattereri

Banded Hognose Snake

Costa, Guedes & Bernils 2021
 
Xenodon severus

Giant Mock Lancehead

Costa, Guedes & Bernils 2021
Genus Xenopholis (South American Flat-headed Snakes)
 
Xenopholis undulatus

Southern Flat-headed Snake

Costa, Guedes & Bernils 2021 Loebmann 2009
 Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.)
Genus Micrurus (American Coral Snakes)
 
Micrurus brasiliensis

Brazilian Coral Snake

Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Pires, Silva, Feitosa, Prudente, Filho & Zaher 2014 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Silva & Sites 1999
 
Micrurus carvalhoi

Brazilian Ribbon Coral Snake

Castro, Oliveira, Gonzalez, Curcio & Feitosa 2017 Costa, Guedes & Bernils 2021 Pires, Silva, Feitosa, Prudente, Filho & Zaher 2014
 
Micrurus hemprichii

Worm-eating Coral Snake

Costa, Guedes & Bernils 2021
 
Micrurus lemniscatus

Northern Ribbon Coral Snake

Costa, Guedes & Bernils 2021
 
Micrurus spixii

Central Amazonian Coral Snake

Costa, Guedes & Bernils 2021 Nascimento, Silva, Feitosa & Prudente 2019
 
Micrurus surinamensis

Amazonian Aquatic Coral Snake

Costa, Guedes & Bernils 2021
 Family Leptotyphlopidae (Thread Snakes)
Genus Trilepida (Three-scaled Thread Snakes)
  *
Trilepida acutirostris

Brazilian Sharp-nosed Thread Snake

Costa, Guedes & Bernils 2021 Francisco, Pinto & Fernandes 2018 Pinto & Curcio 2011 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Trilepida brasiliensis

Bahia Thread Snake

Costa, Guedes & Bernils 2021 Pinto & Curcio 2011 Pinto & Fernandes 2012
 
Trilepida fuliginosa

Preto River Thread Snake

Costa, Guedes & Bernils 2021 Guedes, Queiroz, Sousa & Hamdan 2020 Passos, Caramaschi & Pinto 2006 Pinto & Fernandes 2012
 
Trilepida koppesi

Terenos Thread Snake

Costa, Guedes & Bernils 2021 Filogonio & Canelas 2015 Martins, Koch, Pinto, Folly, Fouquet & Passos 2019 Pinto & Fernandes 2012
 Family Typhlopidae (Typical Blind Snakes)
Genus Amerotyphlops (American Blind Snakes)
 
Amerotyphlops brongersmianus

Bahia Blind Snake

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
 
Amerotyphlops reticulatus

Reticulated Blind Snake

Costa, Guedes & Bernils 2021
 Family Viperidae (Vipers)
Genus Bothrops (American Lanceheads)
 
Bothrops atrox

Common Lancehead

Costa, Guedes & Bernils 2021
 
Bothrops lutzi

Cerrado Lancehead

Carrasco, Grazziotin, Farfan, Koch, Ochoa, Scrocchi, Leynaud & Chaparro 2019 Costa, Guedes & Bernils 2021 Loebmann 2009 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011 Vechio, Prates, Grazziotin, Graboski & Rodrigues 2021
 
Bothrops marmoratus

Marbled Lancehead

Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Silva & Rodrigues 2008
 
Bothrops mattogrossensis

Mato Grosso Lancehead

Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 Silva & Rodrigues 2008
 
Bothrops moojeni

Brazilian Lancehead

Costa, Guedes & Bernils 2021 Recoder, Junior, Camacho, Nunes, Mott, Valdujo, Ghellere, Nogueira & Rodrigues 2011
Genus Crotalus (Typical Rattlesnakes)
 
Crotalus durissus

South American Rattlesnake

Costa, Guedes & Bernils 2021
Genus Lachesis (Bushmasters)
 
Lachesis muta

South American Bushmaster

Costa, Guedes & Bernils 2021

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Chelidae (Austro-American Side-necked Turtles)
Genus Chelus (Matamatas)
 
Chelus fimbriata

Southern Matamata

Costa, Guedes & Bernils 2021 Lima Silveira 2011 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Vargas-Ramirez, Caballero, Morales-Betancourt, Lasso, Amaya, Martinez, Silva Viana, Vogt, Pires Farias, Hrbek, Campbell & Fritz 2020
Genus Mesoclemmys (Hump-backed Toad-headed Turtles)
 
Mesoclemmys gibba

Gibba Toad-headed Turtle

Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
 
Mesoclemmys vanderhaegei

Tobati Toad-headed Turtle

Brito, Strussmann, Kawashita-Ribeiro, Morais, Avila & Campos 2012 Costa, Guedes & Bernils 2021 Maffei, Nascimento, Moya & Donatelli 2016 Marques, Boehm, Brito, Cabrera & Verdade 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
Genus Phrynops (Lesser Toad-headed Turtles)
 
Phrynops geoffroanus

Brazilian Toad-headed Turtle

Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
 Family Geoemydidae (Eastern Pond Turtles)
Genus Rhinoclemmys (Neotropical Wood Turtles)
 
Rhinoclemmys punctularia

Eastern Spotted-legged Turtle

Costa, Guedes & Bernils 2021 Dornas, Malvasio & Pinheiro 2011 Pereira, Santos, Vasconcelos & Oda 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
 Family Kinosternidae (American Mud and Musk Turtles)
Genus Kinosternon (American Mud Turtles)
 
Kinosternon scorpioides

Scorpion Mud Turtle

Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
 Family Podocnemididae (Giant Side-necked Turtles)
Genus Podocnemis (South American Side-necked River Turtles)
 
Podocnemis expansa

Giant South American River Turtle

Costa, Guedes & Bernils 2021 Lubiana & Ferreira-Junior 2009 Portelinha, Malvasio, Pina & Bertoluci 2013 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Salera-Junior, Goncalves Portelinha & Malvasio 2009 Salera-Junior, Malvasio & Goncalves Portelinha 2009
 
Podocnemis unifilis

Yellow-spotted Amazon River Turtle

Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Salera-Junior, Malvasio & Goncalves Portelinha 2009
 Family Testudinidae (Tortoises)
Genus Chelonoidis (South American Tortoises)
 
Chelonoidis carbonarius

South American Red-footed Tortoise

Costa, Guedes & Bernils 2021
 
Chelonoidis denticulatus

South American Yellow-footed Tortoise

Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021